ENTOMOLOGY
Lizards, snails, caterpillars and beetles were common toys to play with when I was a child and enjoyed the countryside in La Palma or at the farm in Tenerife. Later, when I was a teenager I started a collection of beetles, and since then, I haven´t stopped, Entomology having become a Leitmotiv and umbilical cord with nature up to the present. Beetles were probably the hidden reason why I decided to study Biology at the University; when the pressure at home –a mild one– was to do something more “serious”, like Architecture, Agrarian engineering or Philosophy. However, I have never worked in Entomology professionally; but always as a happy and priviledged amateur.
IBCER
When I finished my Biology Degree in La Laguna, the knowledge about Entomology of the Canary Islands was dispersed mainly in foreign institutions and journals. In 1976 I established, along with my wife and my brother, a cultural non-profit private association IBCER (International Bureau of Canarian Entomological Research) with the aim of collecting that information and offering a contact point for all researchers interested in our insect fauna.
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IBCER was more or less useful until the Department of Zoology at the University of La Laguna, and the Natural History Museum of Santa Cruz consolidated their activity in Entomology during the 80,s. In January 1994, IBCER had 53 colleagues registered.
At present, IBCER is dormant, but my library stays open and the same spirit of assistance to any colleague persists in the second floor at home, where I have established my cabinet (you can see a set of photos at the end of this page). Now, I have the intention to provide as much information as I can through this web-site, mainly on Coleoptera, bibliography, and whatever is in my hands, with the only limit being that of available time.
Carabidae
When you are young and looking at bugs, each group has attractive aspects: ants, springtails, spiders, grasshoppers, dragonflies… My interest in ground-beetles goes back to 1969, when I discovered a few black Microlestes close to my golf ball. Its perfect design just fascinated me. The choice was made and it received full support of Don José María Fernández, curator of the Museum of Natural History at Santa Cruz, whom I started to visit regularly.
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Later, in 1975, I presented my Biology Diploma work on the ground-beetles of the laurel forest, and that was the start of a more ambitious project in which I intended to answer questions like: How many species do really live in the Canary Islands? Are they all valid species? Where do they live? It took me about 15 years to perform a full taxonomic and faunal revision to aarive at some satisfactory answers. That was my PhD dissertation (cum laude).
I studied some 23.000 specimens from my own collection as well as from several museum collections (Madrid, Oxford, London, Paris, etc.). A total of 232 species live in the Canary Islands, 70% of which are endemic to the archipelago. I revised all the endemic taxa and described 1 tribe, 4 new genera and 29 new species. In parallel, I established 28 new synonyms. The Monografía de los carábidos de las islas Canarias, was published a year later (1992). It has 734 pages and 313 figures. Some chapters deal with aspects like phenology, dispersal power, and parasites, with some emphasis on biogeography and comments on evolution. But at that time, I did not perform phylogenetic analysis. That came later, but not with carabids.
You can access the Monograph here (PDF) or from the section Publication /Scientific, were all other papers dealing with Carabidae are available.
Laparocerus
Having grown after the monograph on Carabidae, my interest in understanding how species form and why, islands are the best places to study evolution, and beetles the best candidates. I defined a profile for the ideal group: it has to be monophyletic, with as many species as possible, all endemic and without flying capacity. Winner by a wide margin: the genus Laparocerus Schönherr, 1834 (Curculionidae: Entiminae).
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In 1999 I switched from ground-beetles to weevils and designed a project to search for Laparocerus in all of Macaronesia, review the taxonomy, study their ecology and reveal their phylogeny and paths of evolution. If there are any patterns, they should be expressed there.
After 15 years of studying the genus I am still not finished, but getting closer to the end. The Laparocerus are not as rare as previously thought. They are easy to be found by night. I developed efficient techniques for sampling, having collected more than 32.000 specimens, many of them belonging to species previously unknown to science. I prospected the Azores, Madeira and Canary Islands consciously, repeating expeditions to the same island, or to the same place whenever needed. Additionally, I have studied the material kept in all major collections in European museums, plus that sent by colleagues. If you have a recalcitrant species of Laparocerus to be identified, please do not hesitate to write to me.
In the section of Production/Scientific there are several articles about Laparocerus, many describing new species. The number of species level taxa has increased from 123 (when I started) to 215, and there are some more in the pipeline.
I faced the phylogenetic study with the collaboration of Dr. Mariano Hernández Ferrer, from the University Institute of Tropical Diseases and Public Health of the Canary Islands, and hired some technical assistance for laboratory work. For financing part of the costs of the molecular study I received a subvention in 2004-2005 from the Fundación Biodiversidad (Madrid). Needless to say, I had to update my knowledge of Genetics and studied the new methodologies for phylogenetic inference and a set of complex software, all with the good guidance of Mariano. I learned a lot –what I always appreciate– , and I am convinced that he also learned something about beetles. The phylogeny of the Madeiran clade was published in 2008, and that of the Canarian-Moroccan clade should be finished in 2015.
Catalogue of Canarian Coleoptera
Since Coleoptera Atlantidum of Wollaston (1865), by the end of the XX Century, there was no modern account of beetles from the Canary Islands. In collaboration with Dr. Pedro Oromí, from the University of La Laguna, and with the assistance of many colleagues, we published a new catalogue in 2000 (see PDF), updating to 2061 the total number of species and subspecies, of which 1247 are endemic. We also registered 325 recognized synonyms.
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I keep the Coleoptera database alive, and Dr. Oromí is in charge of Entomology in the Banco de datos de Biodiversidad de Canarias (BIOTA), supported by the Canarian Government. Should you have any amendment or contribution to add to the Canarian Coleoptera Database, please feel welcome to contact one of us:
- Dr. Antonio Machado: antonio.machado@telefonica.net
- Dr. Pedro Oromi: poromi@ull.es
When I find a suitable database and some time, it is my intention to include the Canarian Catalogue in this website for online access. For the moment, I keep it in my desktop running on Procite, and oldie, but performing well.
Canarian Entomological Bibliography
One of my main goals at IBCER was to find all the entomological bibliography related to the Canary Islands. That was in the times of cards, typing machines, carbon copies, ordinary letters and smailmail to contact other colleagues, visiting libraries in person, etcetera; something unimaginable for the Internet generation. In 1976 I finished a preliminary list, and in 1987 the book A Bibliography of the Canary Island Entomology was published, which was a milestone.
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Since then, I have been registering all new publications as best as I can. They have grown exponentially, but the new facilities with computers and Internet is a great help. As with the Catalogue of Coleoptera, I hope to be able to transfer my database to an online application within this web-site (probably RefBASE). It is a question of finding the time. Meanwhile, if you need information about any publication, please write to me. In some cases, I can also provide a PDF or scan the original. My library has some 12.000 registers (EndNote X6), and about 4800 are related with Arthropods.
How to get a Collecting Permit for Insects
Collecting insects or any other biological material has become complicated in the Canary Islands. The Spanish national legislation on wildlife conservation and in particular the authorization process for collecting samples was designed with large mammals or birds in the mind, not with insects. To get a permit is possible, but it takes time. Many colleagues just write to me, but I do not extend permits. What I can do is help with the bureaucracy and cross fingers.
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Almost half of the territory of the Canary Islands is covered by some kind of protected area (natural monuments, national parks, rural parks, strict reserves, etc.). To collect in any of these areas, you will need a permit from the conservation authority, which is at present the Island Council (Cabildo Insular) of the respective island, except for national parks (Park Authority). Yes, you will need a permit for each island or national park, if you intend to work on more than one.
Below you will find a link to a new page with application forms, contacts, and some recommendations. Not all Cabildos will ask for the same information, but it refers basically to questions like: what, where, when, how, who, and what for. It will take X time to get a reply, and it is common that the authorization includes some conditions.
If some of your target species are protected because they are threatened, the application procedures are different and the Biodiversity Service of the Autonomous Government will be involved.
SELECTED IMAGES
CABINET WORK
COLLABORATORS
SAMPLING TECHNIQUES
SELECTED VIDEOS
ENTOMOLOGICAL EXPEDITION TO LA GOMERA & EL HIERRO
Director: Antonio Machado Carrillo
Filmed on: Super 8, original footage
© 1971: Antonio Machado Carrillo
'LAPAROCERUS BELLUS' HATCHING OF THE EGG
Director: Antonio Machado Carrillo
Location: Tenerife, Canary Islands
© 2020: Antonio Machado Carrillo
'LAPAROCERUS RUTERI' HATCHING OF THE EGG
Director: Antonio Machado Carrillo
Location: Tenerife, Canary Islands
© 2020: Antonio Machado Carrillo
MORE INFO
LIST OF NEW TAXA DESCRIBED
Tribes
- Dicrodontini Mach. 1992, Carabidae: Zuphiinae
Genera
- Canarobius Mach. 1987, Carabidae, Trechinae, Trechodini
- Spelaeovulcania Mach. 1987, Carabidae, Trechinae, Trechodini
- Wolltinerfia Mach. 1985 nom.nov. pro Wollastonia Mach. 1984, Carabidae Pterostichinae
- Orzolina Mach. 1987c, Carabidae, Trechinae, Bembidini
Subgenera
- Gietopus 1992, del género Wolltinerfia Mach. 1985, Carabidae
- Lauricalathus 1992, del género Calathus Bon., Carabidae
- Fortunotrox 2011, del género Laparocerus Schoenh. 1834, Curculionidae
- Pseudatlantis 2008, del género Laparocerus Schoenh. 1834, Curculionidae
- Wollastonius 2008, del género Laparocerus Schoenh. 1834, Curculionidae
- Atlantodes 2008, del género Laparocerus Schoenh. 1834, Curculionidae
- Faycanius Mach. 2012, del género Laparocerus Schoenh. 1834, Curculionidae
- Aridotrox , 2017, del género Laparocerus Schoenh. 1834, Curculionidae
- Belicarius , 2017, del género Laparocerus Schoenh. 1834, Curculionidae
- Bencomius, 2017, del género Laparocerus Schoenh. 1834, Curculionidae
- Canariotrox , 2017, del género Laparocerus Schoenh. 1834, Curculionidae
- Purpuranius, 2017, del género Laparocerus Schoenh. 1834, Curculionidae
Species
- Pimelia fernandezlopezi Mach. 1979, Tenebrionidae.
- Canarobius chusyae Mach. 1988. Carabidae
- Canarobius oromii Mach. 1988. Carabidae
- Thalassophilus subterraneus Mach. 1989. Carabidae
- Spelaeovulcania canariensis Mach. 1987. Carabidae
- Trechus minioculatus Mach. 1987. Carabidae
- Trechus benahoaritus Mach. 1989. Carabidae
- Lymnastis subovatus Mach. 1992. Carabidae
- Lymnastis thoracicus Mach. 1992. Carabidae
- Tachys fortunatus Mach. 1989. Carabidae
- Orzolina thalassophila Mach. 1987. Carabidae
- Wolltinerfia tenerifae (Mach. 1984). Carabidae (sub. Wollastonia)
- Wolltinerfia martini (Mach. 1984). Carabidae (sub. Eutrichopus)
- Calathus refleximargo Mach. 1992. Carabidae
- Calathus pilosipennis Mach. 1992. Carabidae
- Calathidius brevithorax Mach. 1992. Carabidae
- Licinopsis angustula Mach. 1987. Carabidae
- Licinopsis schurmanni Mach. 1987. Carabidae (como ssp. de L. angustula)
- Zabrus canaricus Mach. 1992. Carabidae
- Bradycellus feloi Mach. 1992. Carabidae [= syn.]
- Paradromius iucundus Mach. 1992. Carabidae
- Paradromius tamaranus Mach. 1992. Carabidae
- Paradromius amplius Mach. 1992. Carabidae
- Paradromius exornatus Mach. 1992. Carabidae
- Paradromius purpurarius Mach. 1992. Carabidae
- Paradromius hariensis Mach. 1992. Carabidae
- Paradromius scholzi Mach. 1992. Carabidae
- Philorhizus mateui Mach. 1992. Carabidae
- Philorhizus franzi Mach. 1992. Carabidae
- Metadromius palmi Mach. 1992. Carabidae
- Parazuphium feloi Mach. 1998. Carabidae
- Drouetius borgesi Mach. 2009. Curculionidae
- Drouetius oceanicus Mach. 2009. Curculionidae
- Laparocerus bacalladoi Mach. 2005. Curculionidae
- Laparocerus junonius Mach., 2007. Curculionidae
- Laparocerus aguiari Mach. 2007. Curculionidae
- Laparocerus subparalellus Mach., 2007. Curculionidae
- Laparocerus boticarius Mach., 2007. Curculionidae
- Laparocerus roudieri Mach., 2007. Curculionidae
- Laparocerus chasnensis Mach.,2007. Curculionidae
- Laparocerus tinguaro Mach., 2007. Curculionidae
- Laparocerus humeralis Mach., 2007. Curculionidae
- Laparocerus acutipennis Mach., 2007. Curculionidae
- Laparocerus exiguus Mach., 2007. Curculionidae
- Laparocerus anopthalmus Mach., 2007. Curculionidae
- Laparocerus hupalupa Mach., 2007. Curculionidae
- Laparocerus orone Mach., 2007. Curculionidae
- Laparocerus oculatissimmus Mach., 2007. Curculionidae
- Laparocerus depressus Mach., 2007. Curculionidae
- Laparocerus spinimanus Mach., 2007. Curculionidae
- Laparocerus mulagua Mach., 2007. Curculionidae
- Laparocerus benchijigua Mach., 2007. Curculionidae
- Laparocerus inermis Mach., 2007. Curculionidae
- Laparocerus heres Mach., 2007. Curculionidae
- Laparocerus madeirensis, Mach. 2008. Curculionidae
- Laparocerus edaphicus Mach. 2008. Curculionidae
- Laparocerus prainha Mach. 2008. Curculionidae
- Laparocerus oromii Mach. 2008. Curculionidae
- Laparocerus stuebeni Mach. 2008. Curculionidae
- Laparoceerus hobbit Mach. 2008. Curculionidae
- Laparocerus cryptus Mach. 2008. Curculionidae
- Laparocerus silvaticus Mach. 2008. Curculionidae
- Laparocerus serrado Mach. 2008. Curculionidae
- Laparocerus lopezi Mach. 2008. Curculionidae
- Laparocerus aeneotincutus Mach. 2009. Curculionidae
- Laparocerus elongatus Mach. 2009. Curculionidae
- Laparocerus supranubius Mach. 2009. Curculionidae
- Laparocerus astralis Mach. 2009. Curculionidae
- Laparocerus amicorum Mach. 2009. Curculionidae
- Laparocerus cristatus Mach. 2009. Curculionidae
- Laparocerus feloi Mach. 2009. Curculionidae
- Laparocerus morrisi Mach. 2009. Curculionidae
- Laparocerus decipiens Mach. 2009. Curculionidae
- Laparocerus tanausu Mach. 2009. Curculionidae
- Laparocerus arrochai Mach. 2009. Curculionidae
- Laparocerus acyphus Mach. 2009. Curculionidae
- Laparocerus femoralis Mach. 2009. Curculionidae
- Laparocerus mucronatus Mach. 2009. Curculionidae
- Laparocerus tarsalis Mach. 2009. Curculionidae
- Laparocerus cephalotes Mach. 2011. Curculionidae
- Laparocerus bimbache Mach. 2011. Curculionidae
- Laparocerus cavernarius Mach. 2011. Curculionidae
- Laparocerus hypogeus Mach. 2011. Curculionidae
- Laparocerus rotundatus Mach. 2011. Curculionidae
- Laparocerus pilosiventris Mach. 2011. Curculionidae
- Laparocerus confusus Mach. 2011. Curculionidae
- Laparocerus juelensis Mach. 2011. Curculionidae
- Laparocerus magnificus Mach. 2011. Curculionidae
- Laparocerus iruene Mach. & García 2011. Curculionidae
- Tarphius jandiensis Mach. 2011. Zopheridae
- Tarphius desertaensis Mach. 2011. Zopheridae
- Laparocerus maxorata Mach. 2011. Curculionidae
- Laparocerus colonnellii Mach. 2011. Curculionidae
- Laparocerus calvus Mach. 2011. Curculionidae
- Laparocerus xericola Mach. 2011. Curculionidae
- Laparocerus fraterculus Mach. 2011. Curculionidae
- Laparocerus longipennis Mach. 2011. Curculionidae
- Laparocerus estevezi Mach. 2012. Curculionidae
- Laparocerus arcanus Mach. 2012. Curculionidae
- Laparocerus dissidens Mach. 2012. Curculionidae
- Laparocerus bentejui Mach. 2012. Curculionidae
- Laparocerus anniversarius Mach. 2012. Curculionidae
- Laparocerus franzi Mach. 2012. Curculionidae
- Laparocerus teldense Mach. 2012. Curculionidae
- Laparocerus fraudulentus Mach. 2012. Curculionidae
- Laparocerus hytricoides Mach. 2012. Curculionidae
- Laparocerus rugosivertex Mach. 2012. Curculionidae
- Laparocerus ornatus Mach. 2012. Curculionidae
- Laparocerus marmoratus Mach. 2012. Curculionidae
- Laparocerus tirmensis Mach. 2012. Curculionidae
- Laparocerus semipilosus Mach. 2012. Curculionidae
- Laparocerus osorio Mach. 2012. Curculionidae
- Laparocerus tirajana Mach. 2012. Curculionidae
- Oromia thoracica Mach. & López, 2015. Curculionidae
- Laparocerus merigensis Mach. 2015. Curculionidae
- Laparocerus dilutus Mach., 2015. Curculionidae
- Laparocerus notatus Mach., 2015. Curculionidae
- Laparocerus teselinde Mach., 2015. Curculionidae
- Laparocerus quadratus Mach., 2015. Curculionidae
- Laparocerus macilentus Mach., 2015. Curculionidae
- Laparocerus sonchiphagus Mach., 2015. Curculionidae
- Laparocerus tenicola Mach., 2015. Curculionidae
- Laparocerus anagae Mach., 2015. Curculionidae
- Laparocerus campestris Mach., 2015. Curculionidae
- Laparocerus abona Mach., 2016. Curculionidae
- Laparocerus tauce Mach., 2016. Curculionidae
- Laparocerus crotchi Mach., 2016. Curculionidae
- Laparocerus gerodes Mach, 2016. Curculionidae
- Laparocerus tafadensis Mach., 2016. Curculionidae
- Laparocerus soniae Mach., 2016. Curculionidae
- Laparocerus auarita Mach., 2016. Curculionidae
- Laparocerus canescens Mach., 2016. Curculionidae
- Laparocerus punctiger Mach., 2016. Curculionidae
- Laparocerus sanctaecrucis Mach,, 2016. Curculionidae
- Laparocerus federico Mach. 2018. Curculionidae
- Herpisticus famarae Mach. 2020. Curculionidae
- Herpisticus betancuriae Mach. 2020. Curculionidae
- Herpisticus jandiensis Mach. 2020. Curculionidae
- Herpisticus rectipes Mach. 2020. Curculionidae
- Herpisticus guanarteme Mach. 2020. Curculionidae
- Herpisticus denudatus Mach. 2020. Curculionidae
- Herpisticus gigas Mach. 2020. Curculionidae
- Herpisticus guayarmina Mach. 2020. Curculionidae
- Herpisticus tasarticus Mach. 2020. Curculionidae
- Herpisticus scopulus Mach. 2020. Curculionidae
- Herpisticus nanus Mach. 2020. Curculionidae
- Herpisticus daute Mach. 2020. Curculionidae
- Herpisticus aridicola Mach. 2020. Curculionidae
- Herpisticus gomerensis Mach. 2020. Curculionidae
- Herpisticus hispidus Mach. 2020. Curculionidae
- Herpisticus bobadillae Mach. 2020. Curculionidae
Subspecies
- Aepus gracilicornis meridionalis Mach. 1992. Carabidae
- Olisthopus glabratus tamaranus Mach. 1992. Carabidae
- Platyderus alticola hierroensis Mach. 1992. Carabidae
- Platyderys alticola gomeraensis Mach. 1992. Carabidae
- Licinopsis obliterata franzi Mach. 1987. Carabidae
- Licinopsis picescens gomerita Mach. 1987. Carabidae
- Paradromius iucundus gillerforsi Mach. 1992. Carabidae
- Paradromius exornatus furvus Mach. 1992. Carabidae
- Paradromius strigifrons grancanariensis Mach. 1992. Carabidae
- Paradromius insularis illautus Mach. 1992. Carabidae
- Dromius angustus dissimilis Mach. 1992. Carabidae
- Philorhizus atlanticus palmensis Mach. 1992. Carabidae
- Philorhizus incertus jandiensis Mach. 1992. Carabidae
- Dicrodontus brunneus exilis Mach. 1992. Carabidae
- Parazuphium damascenum canariense Mach. 1992. Carabidae
- Drouetius azoricus nitens Mach. 2009. Curculionidae
- Drouetius azoricus parallelirostris Mach. 2009. Curculionidae
- Drouetius azoricus separandus Mach. 2009. Curculionidae
- Drouetius borgesi centralis Mach. 2009. Curculionidae
- Drouetius borgesi sanctimichaelis Mach. 2009. Curculionidae
- Drouetius oceanicus tristis Mach. 2009. Curculionidae
- Laparocerus aethiops garajonay Mach., 2007. Curculionidae
- Laparocerus heres jocoensis Mach., 2007. Curculionidae
- Laparocerus mateui tuberosus Mach. 2011. Curculionidae
- Laparocerus orone aridane Mach. 2009. Curculionidae
- Laparocerus orone hierroensis Mach. 2011. Curculionidae
- Laparocerus zarazagai subreflexus Mach. & García 2011. Curculionidae
- Laparocerus rasus jandiensis Mach. 2011. Curculionidae
- Laparocerus rasus betancor Mach. 2011. Curculionidae
- Laparocerus curvipes famara Mach. 2011. Curculionidae
- Laparocerus garretai albosquamosus Mach. 2011. Curculionidae
- Laparocerus susicus inexpectatus Mach. 2011. Curculionidae
- Laparocerus susicus montanus Mach. 2011. Curculionidae
- Laparocerus squamosus tasarticus Mach. 2012. Curculionidae
- Laparocerus alluaudi aytamis Mach. 2012. Curculionidae
- Laparocerus bentejui delicatulus Mach. 2012. Curculionidae
- Laparocerus bentejui robustus Mach. 2012. Curculionidae
- Laparocerus subnodosus montivagans Mach. 2013. Curculionidae
- Laparocerus sanchezi arures , 2016. Curculionidae
- Laparocerus obscurus daute , 2016. Curculionidae
- Laparocerus tinguaro tabornoi , 2016. Curculionidae
- Laparocerus dissimilis alticola , 2016. Curculionidae
- Laparocerus dissimilis infernalis , 2016. Curculionidae
- Laparocerus tenuepunctatus oppositus , 2016. Curculionidae
- Herpisticus subvestitus pseudolanatus Mach. 2020. Curculionidae
- Herpisticus hierrensis benahoare Mach. 2020. Curculionidae
PRELIMINARY CATALOGUE OF LAPAROCERUS (UPDATED MAY 2017)
Subfamily Entiminae Schoenherr, 1823
Tribe Laparocerini Lacordaire, 1863
Genus Laparocerus Schoenherr, 1834: 530 type species Laparocerus morio Boheman, 1834
= Cyphoscelis Wollaston, 1854: 356 type species Cyphoscelis distorta Wollaston, 1854
Subgenus Amyntas Wollaston, 1865: Appendix, 55 type species Lichenophagus incomptus Wollaston, 1865
= Canopus Wollaston, 1864: 337 [HN] type species Atlantis subnebulosa Wollaston, 1864
arrochai Machado, 2009: 190 N: CI (La Palma)
bellus Roudier, 1957a: 31 N: CI (Tenerife)
cristatus Machado, 2009: 193 N: CI (La Palma)
fernandezi Roudier, 1957a: 34 N: CI (Tenerife)
incomptus Wollaston, 1865: Appendix, 55 (Lichenophagus) N: CI (El Hierro)
ruteri Roudier, 1957a: 33 N: CI (Tenerife)
subnebulosus Wollaston, 1864: 337 (Atlantis) N: CI (Gran Canaria)
tanausu Machado, 2009: 195 N: CI (La Palma)
tetricus Boheman, 1834: 542 (Eremnus) N: CI (Tenerife)
= simplex Brullé, 1839: 71 (Otiorhynchus)
tibialis Wollaston, 1864: 338 (Atlantis) N: CI (Tenerife)
uyttenboogaarti Zumpt, 1940: 677 N: CI (Tenerife)
Subgenus Anillobius Fauvel, 1907: 71 type species Anillobius solifuga Fauvel, 1907
solifuga Fauvel, 1907: 72 (Anillobius) N: MR (Madeira)
portosantoi Franz 1970: 3 (Anillobius) N: MR (Porto Santo)
Subgenus Aridotrox Machado 2017: 33 type species Laparocerus rasus Wollaston, 1864
colonnellii Machado, 2011b: 232 N: CI (Fuerteventura)
dispar Wollaston, 1864: 357 N: CI (Lanzarote)
rasus betancor Machado, 2011b: 223 N: CI (Fuerteventura)
rasus jandiensis Machado, 2011b: 224 N: CI (Fuerteventura)
rasus rasus Wollaston, 1864: 354 N: CI (Lanzarote)
susicus inexpectatus Machado, 2011b: 229 N: MO
tessellatus Hustache, 1932: 43 (Cyclomaurus) [HN]
susicus montanus Machado, 2011b: 229 N: MO
susicus susicus Escalera, 1914: 424 (Cyclobarus) N: MO
xericola Machado, 2011b: 231 N: CI (Fuerteventura)
Subgenus Atlantis Wollaston, 1854: 361 type Atlantis lamellipes Wollaston, 1854
calcatrix Wollaston, 1854: 366 (Atlantis) N: MR (Madeira)
= calcator Marseul, 1872: 546
clavatus Wollaston, 1854: 363 (Atlantis) N: MR (Madeira)
hobbit Machado, 2008a: 318 N: MR (Madeira)
lamellipes Wollaston, 1854: 364 (Atlantis) N: MR (Madeira)
lauripotens Wollaston, 1854: 369 (Atlantis) N: MR (Madeira)
= australis Wollaston, 1864: 370 (Atlantis)
lindbergi Roudier, 1963: 133 N: MR (Madeira)
madeirensis Machado, 2008a: 316 N: MR (Madeira)
noctivagans Wollaston, 1854: 367 (Atlantis) [NP] N: MR (Madeira)
= piceus Boheman, 1834: 531 [NO]
serrado Machado, 2008a: 319 N: MR (Madeira)
undulatus Wollaston, 1862: 289 N: MR (Madeira)
vespertinus Wollaston, 1854: 371 (Atlantis) N: MR (Madeira)
Subgenus Atlantodes Machado, 2008a: 312 type species Atlantis mendax Wollaston, 1854
colasi Roudier, 1958: 205 N: MR (Madeira)
inconstans Wollaston, 1854: 375 (Atlantis) N: MR (Porto Santo)
instabilis Wollaston, 1854: 377 (Atlantis) N: MR (Porto Santo)
lanatus Wollaston, 1854: 372 (Atlantis) N: MR (Madeira)
mendax Wollaston, 1854: 376 (Atlantis) N: MR (Porto Santo)
navicularis Wollaston, 1854: 374 (Atlantis) N: MR (Porto Santo)
prainha Machado, 2008a: 323 N: MR (Madeira)
Subgenus Belicarius Machado 2017: 35 type species Laparocerus mendicus Wollaston, 1864
bentejui bentejui Machado, 2012: 7 N: CI (Gran Canaria)
bentejui delicatulus Machado, 2012: 8 N: CI (Gran Canaria)
bentejui robustus Machado, 2012: 10 N: CI (Gran Canaria)
exiguus Machado, 2007b: 11 N: CI (La Gomera)
exophthalmus Machado, 2007b: 12 N: CI (La Gomera)
feloi Machado, 2009: 203 N: CI (La Palma)
longiclava Lindberg, 1953: 15 N: CI (Tenerife)
magnificus Machado, 2011a: 70 N: CI (La Gomera)
mateui mateui Roudier, 1954: 85 N: CI (La Gomera)
mateui tuberosus Machado, 2011a: 74 N: CI (El Hierro)
mendicus Wollaston, 1864: 355 N: CI (El Hierro)
morrisi Machado, 2009: 218 N: CI (La Palma)
oculatissimus Machado, 2007b: 14 N: CI (La Gomera)
rotundatus Machado, 2011a: 65 N: CI (La Gomera)
sanchezi arures Machado, 2016: 33 N: CI (La Gomera)
sanchezi sanchezi Roudier, 1957a: 24 N: CI (El Hierro)
= nitidirostris Roudier, 1957a: 25
Subgenus Bencomius Machado 2017: 35 type species Laparocerus grossepunctatus Wollaston, 1864
bolivari Uyttenboogaart, 1937: 97 N: CI (Tenerife)
boticarius Machado, 2007a: 39 N: CI (Tenerife)
combrecitensis Roudier, 1957c: 90 [RNJ N: Cl (La Palma)
palmensis Roudier, 1957a: 20 [HN]
crassifrons Wollaston, 1864: 345 N: CI (Tenerife)
edaphicus Machado, 2008b: 1278 N: CI (Tenerife)
escaleraorum Uyttenboogaart, 1937: 96 N: CI (Tenerife)
gomerensis Lindberg, 1953: 15 N: CI (La Gomera)
grossepunctatus Wollaston, 1864: 344 N: CI (Tenerife)
scapularis Wollaston, 1864: 347 N: CI (Tenerife)
= wollastoni Marseul, 1872: 521
subparallelus Machado, 2007a: 43 N: CI (Tenerife)
supranubius Machado, 2009: 210 N: CI (La Palma)
tenuepunctatus oppositus Machado, 2016: 16 N: CI (Tenerife)
tenuepunctatus tenuepunctatus Roudier, 1957a: 20 N: CI (Tenerife)
undatus Wollaston, 1864: 342 N: CI (Tenerife)
Subgenus Canariotrox Machado 2017: 35 type species Laparocerus inaequalis Wollaston, 1864
abona Machado, 2016: 14 N: CI (Tenerife)
acyphus Machado, 2009: 189 N: CI (La Palma)
aeneotinctus Machado, 2009: 185 N: CI (La Palma)
affinis Wollaston, 1864: 358 N: CI
aguiari Machado, 2007a: 36 N: CI (Tenerife)
crassus Roudier, 1957a: 28 N: CI (Tenerife)
estevezi Machado, 2012: 13N: CI (Gran Canaria)
femoralis Machado, 2009: 188 N: CI (La Palma)
hirtus Wollaston, 1864: 348 N: CI (Gran Canaria)
= doramasensis Uyttenboogart, 1929: 352
inaequalis globulipennis Wollaston, 1864: 349 N: CI (La Palma)
= globosipennis Marseul, 1872: 531
inaequalis inaequalis Wollaston, 1863: 220 N: CI (Tenerife)
occidentalis Wollaston, 1864: 350 N: CI (El Hierro)
rugosicollis Uyttenboogaart, 1937: 98 N: CI (Tenerife)
tauce Machado, 2016: 16 N: CI (Tenerife)
vestitus Wollaston 1864: 358 N: CI
= velatus Marseul, 1872: 532
Subgenus Faycanius Machado, 2012: 29 type species Otiorhynchus squamosus Brullé, 1839
alluaudi alluaudi Uyttenboogaart, 1940: 55 N: CI (Gran Canaria)
alluaudi aytamis Machado, 2012: 33 N: CI (Gran Canaria)
anniversarius Machado, 2012: 37 N: CI (Gran Canaria)
arcanus Machado, 2012: 32 N: CI (Gran Canaria)
dissidens Machado, 2012: 35 N: CI (Gran Canaria)
squamosus squamosus Brullé, 1839: 71 (Otiorhynchus) N: CI (Gran Canaria)
= seniculus Wollaston, 1864: 353
squamosus tasarticus Machado, 2012: 21 N: CI (Gran Canaria)
Subgenus Fernandezius Roudier, 1957a: 40 type species Lichenophagus tesserula Wollaston, 1864
anagae Machado, 2015: 126 N: CI (Tenerife)
campestris Machado, 2015: 132 N: CI (Tenerife)
impressicollis Wollaston, 1864: 368 (Lichenophagus) N: CI (Tenerife)
macilentus Machado, 2015: 125 N: CI (Tenerife)
persimilis Wollaston, 1864: 365 (Lichenophagus) N: CI (Tenerife)
sculptipennis montivagans Machado 2013: 69 N: CI (La Palma)
= palmensis Roudier 1957a: 43 (Lichenophagus) [HN]
sculptipennis sculptipennis Wollaston, 1864: 367 (Lichenophagus) N: CI (La Palma)
seriesetosus Wollaston, 1864: 365 (Lichenophagus) N: CI [La Palma]
sonchiphagus Machado, 2015: 120 N: CI (Tenerife)
subcalvus Wollaston, 1864: 336 (Lichenophagus) N: CI (El Hierro)
subnodosus Wollaston, 1864: 336 (Lichenophagus) N: CI (Tenerife)
tenicola Machado, 2015: 128 N: CI (Tenerife)
tesserula Wollaston, 1864: 364 (Lichenophagus) N: CI (Tenerife)
Subgenus Fortunotrox Machado, 2011a: 58 type species Laparocerus gracilis Wollaston, 1865
acutipennis Machado, 2007b: 5 N: CI (La Gomera)
benchijigua Machado, 2007b: 6 N: CI (La Gomera)
confusus Machado, 2011a: 62 N: CI (La Gomera)
crotchi Machado, 2016: 30 N: CI (La Gomera)
decipiens Machado, 2009: 214 N: CI (La Palma)
depressus Machado, 2007b: 7 N: CI (La Gomera)
gracilis Wollaston, 1864: 356 N: CI (La Gomera)
= gomeraecola Marseul, 1872: 531
indutus Wollaston, 1865 (app.): 53 N: CI (La Gomera)
tectus Marseul, 1872: 535
juelensis Machado, 2011a: 64 N: CI (La Gomera)
orone aridane Machado, 2009: 211 N: CI (La Palma)
orone hierrensis Machado, 2011a: 80 N: CI (El Hierro)
orone orone Machado, 2007b: 3 N: CI (La Gomera)
puncticollis Wollaston, 1864: 362 N: CI (El Hierro)
spinimanus Machado, 2007b: 9 N: CI (La Gomera)
subopacus Wollaston, 1865 (app.): 52 N: CI (La Gomera)
Subgenus Guanchotrox Alonso-Zarazaga & Lyal, 1999: 11 [RN] type species Laparocerus canariensis Boheman, 1843
= Amphora Wollaston, 1864: 335 [HN] type species Laparocerus canariensis Boheman, 1843
amicorum Machado, 2009: 212 N: CI (La Palma)
astralis Machado, 2009: 206 N: CI (La Palma)
buenavistae Roudier, 1957a: 29 N: CI (Tenerife)
canariensis Boheman, 1842: 228 N: CI (Tenerife)
cephalotes Machado, 2011a: 78 N: CI (El Hierro)
debilis Wollaston, 1865: Appendix, 53 N: CI (Tenerife)
dissimilis alticola Machado, 2016: 22 N: CI (Tenerife)
dissimilis infernalis Machado 2016: 23 N: CI (Tenerife)
dissimilis dissimilis Lindberg, 1950: 17 N: CI (Tenerife)
elongatus Machado, 2009: 198 N: CI (La Palma)
gerodes Machado, 2016: 23 N: CI (La Gomera)
humeralis Machado, 2007b: 23 N: CI (La Gomera)
junonius Machado, 2007b: 19 N: CI (La Gomera)
marmoratus Machado, 2012: 22 N: CI (Gran Canaria)
mucronatus Machado, 2009: 201 N: CI (La Palma)
obscurus daute Machado, 2016: 21 N: CI (Tenerife)
obscurus obscurus Wollaston, 1864: 355 N: CI (Tenerife)
= obscurellus Marseul, 1872: 539
obtriangularis Wollaston, 1864: 351 N: CI (Tenerife)
ornatus Machado, 2012:20 N: CI (Gran Canaria)
palmensis Lindberg, 1953: 16 N: CI (La Palma)
seminitens Lindberg, 1950: 17 N: CI (Tenerife)
tafadensis Machado, 2016: 28 N: CI (Tenerife)
tarsalis Machado, 2009: 205 N: CI (La Palma)
tenellus Wollaston, 1864: 362 N: CI (Tenerife)
tirmensis Machado, 2012: 21 N: CI (Gran Canaria)
roudieri Machado, 2007b: 22 N: CI (La Gomera)
tinguaro tabornoi Machado, 2016: 29 N: CI (Tenerife)
tinguaro tinguaro Machado, 2007a: 44 N: CI (Tenerife)
transversus Lindberg, 1950: 16 N: CI (Tenerife)
Subgenus Laparocerus Schoenherr, 1834: 530 type species Laparocerus morio Boheman, 1834
= Cyphoscelis Wollaston, 1854: 356 type species Cyphoscelis distorta Wollaston, 1854
chaoensis cevadae Roudier, 1961: 62 N: MR (Ilheu do Desembarcadouro)
chaoensis chaoensis Uyttengoogaart, 1940: 57 N: MR (Ilheu de Chão)
= vandeli Roudier 1958: 199
cryptus Machado, 2008a: 315 N: MR
distortus Wollaston, 1854: 357 (Cyphoscelis) N: MR (Madeira)
morio Boheman, 1834: 531 N: MR (Madeira)
= obesulus Desbrochers, 1904: 63
Subgenus Lichenophagus Wollaston, 1854: 389 type species Lichenophagus fritillus Wollaston, 1854
acuminatus Wollaston, 1854: 391 E: MR (Porto Santo)
fritillus Wollaston, 1854: 390 E: MR (Deserta Grande)
Subgenus Machadotrox Alonso-Zarazaga & Lyal, 1999: 10 type species Otiorhynchus sculptus Brullé, 1839
aethiops aethiops Wollaston, 1864: 347 N: CI (El Hierro)
aethiops garajonay Machado, 2007b: 15 N: CI (La Gomera)
cavernarius Machado, 2011a: 84 N: CI (El Hierro)
chasnensis Machado, 2007a: 41 N: CI (Tenerife)
dacilae García, 1988: 45 N: CI (La Palma)
excavatus excavatus Wollaston, 1863: 219 N: CI (Tenerife)
excavatus lugubris Wollaston, 1864: 341 N: CI (Tenerife)
hupalupa Machado, 2007b: 16 N: CI (La Gomera)
hypogeus Machado, 2011a: 82 N: CI (El Hierro)
idafe García & Alonso-Zarazaga, 2011: 73 N: CI (La Palma)
inermis Machado, 2007b: 18 N: CI (La Gomera)
iruene Machado & García, 2010: 65 N: CI (La Palma)
laevis Roudier, 1957a: 18 N: CI (La Palma)
= politus Roudier, 1957a: 19
oromii Machado, 2008b: 1282 N: CI (La Gomera)
sculptus Brullé, 1839: 71 (Otiorhynchus) N: CI (La Palma)
zarazagai subreflexus Machado & García, 2010: 68 N: CI (La Palma)
zarazagai zarazagai García & Oromí, 1997: 154 N: CI (La Palma)
Subgenus Mateuius Roudier, 1957a: 40 type species Lichenophagus auctus Wollaston, 1864
amplificatus Wollaston, 1865: 330 (Lichenophagus) N: CI (La Gomera)
auctus Wollaston, 1864: 363 (Lichenophagus) N: CI (El Hierro)
buccatrix Wollaston, 1865: Appendix, 54 (Lichenophagus) N: CI (La Gomera)
dilutus Machado, 2015: 105 N: CI (La Gomera)
merigensis Machado, 2015: 108 N: CI (La Gomera)
notatus Machado, 2015: 105 N: CI (La Gomera)
quadratus Machado, 2015: 108 N: CI (La Gomera)
teselinde Machado 2015: 106 N: CI (La Gomera)
Subgenus Pecoudius Roudier, 1957a type species Laparocerus cymbula Roudier, 1957a (= Cyphoscelis eliasenae Uyttenboogaart, 1929)
compactus Wollaston, 1864: 359 N: CI (Gran Canaria)
eliasenae Uyttenboogaart, 1929: 353 (Cyphoscelis) N: CI (Gran Canaria)
= cymbula Roudier, 1957a: 38
franzi Machado, 2012: 10 N: CI (Gran Canaria)
sulcirostris Wollaston, 1864: 359 N: CI (Gran Canaria)
= canalirostris Marseul, 1872: 537
teldensis Machado, 2012: 11 N: CI (Gran Canaria)
[Pecoudius s.l.]
auarita Machado, 2016: 9 N: CI (La Palma)
bimbache Machado, 2011a: 74 N: CI (El Hierro)
brunneus Lindberg, 1953: 14 N: CI (Gran Canaria)
canescens Machado, 2016: 8 N: CI (Tenerife)
crassirostris Wollaston, 1864: 345 N: CI (Gran Canaria)
fraudulentus Machado, 2012: 14 N: CI (Gran Canaria)
freyi Uyttenboogaart, 1940: 675 N: CI (Tenerife)
garretai albosquamosus Machado, 2011b: 237 N: SG (Gran Pitón)
garretai garretai Uyttenboogaart, 1940: 56 N: SG (Selvagem Grande)
grayanus Wollaston, 1865: 309 (Atlantis) [RN] N: CI (Gran Canaria)
= angustulus Wollaston, 1863: 219 (Atlantis) [HN]
hystricoides Machado, 2012: 16 N: CI (Gran Canaria)
inconspectus Roudier, 1957a: 27 N: CI (Gran Canaria)
lepidopterus lepidopterus Wollaston, 1864: 352 N: CI
= bellopterus Marseul, 1872: 527
lepidopterus pecoudi Roudier, 1957a: 22 N: CI (Gran Canaria)
lopezi Machado, 2008b: 1285 N: CI (Gran Canaria)
microphthalmus Lindberg, 1950: 16 N: CI (Gran Canaria)
mulagua Machado, 2007b: 25 N: CI (La Gomera)
obsitus Wollaston, 1864: 361 N: CI (Gran Canaria)
= persitus Marseul, 1872: 534
osorio Machado, 2012: 25 N: CI (Gran Canaria)
propinquus Lindberg, 1953: 14 N: CI (Gran Canaria)
punctiger Machado, 2016: 6 N: CI (Tenerife)
rugosivertex Machado, 2012: 18 N: CI (Gran Canaria)
semipilosus Machado, 2012: 23 N: CI (Gran Canaria)
separandus Lindberg, 1953: 14 N: CI (Gran Canaria)
soniae Machado, 2016: 35 N: CI (Gran Canaria)
tessellatus Brullé, 1839: 72 (Omias) N: CI (Tenerife)
tirajana Machado, 2012: 28 N: CI (Gran Canaria)
vicinus Lindberg, 1953: 15 N: CI (Gran Canaria)
Subgenus Pseudatlantis Machado, 2008a: 312 type species Atlantis excelsa Wollaston, 1854
abditus Roudier, 1963: 136 N: MR (Madeira)
excelsus Wollaston, 1854: 378 (Atlantis) N: MR (Madeira)
schaumii Wollaston, 1854: 379 (Atlantis) N: MR
= forae Wollaston, 1854: 380 (Atlantis)
silvaticus Machado, 2008a: 319 N: MR (Madeira)
stuebeni Machado, 2008a: 321 N: MR (Madeira)
Subgenus Purpuranius Machado 2017: 33 type species Laparocerus maxorata Machado, 2011
calvus Machado, 2011b: 215 N: CI (Fuerteventura)
curvipes curvipes Lindberg, 1950: 17 N: CI (Tenerife)
curvipes espanoli Roudier, 1954: 84 N: CI (Fuerteventura, Lanzarote)
curvipes famarae Machado, 2011b: 219 N: CI (Lanzarote)
fraterculus Machado, 2011b: 211 N: CI (Fuerteventura)
longipennis Machado, 2011b: 213 N: CI (Fuerteventura)
maxorata Machado, 2011b: 208 N: CI (Fuerteventura)
Subgenus Wollastonius Machado, 2008a: 313 type species Omias ventrosus Wollaston, 1854
aenescens Wollaston, 1854: 383 (Omias) N: MR (Madeira)
angustulus Wollaston, 1857: 116 (Omias) N: MR (Madeira)
ventrosus Wollaston, 1854: 382 (Omias) N: MR (Madeira)
waterhousei Wollaston, 1854: 384 (Omias) N: MR
Genus Laparocerus, species incertae sedis
bacalladoi Machado, 2005: 540 N: CI (Tenerife)
depilis Roudier, 1957a: 22 N: CI (Tenerife)
ellipticus Wollaston, 1863: 220 N: CI
heres heres Machado, 2007b: 29 N: CI (La Gomera)
heres jocoensis Machado, 2007b: 35 N: CI (Tenerife)
inflatus Wollaston, 1865: Appendix, 41 N: CI (La Gomera)
= tumens Marseul, 1872: 526
pilosiventris Machado, 2011a: 68 N: CI (La Gomera)
sanctaecrucis Machado, 2016: 11 N: CI (Tenerife)
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Wollaston T. V. 1862: On additions to the Madeiran Coleoptera. Annals and Magazine of Natural History (3) 10: 287-293.
Wollaston T. V. 1863: Diagnostic notices of new Canarian Coleoptera. Annals and Magazine of natural History (3) 11: 214-221.
Wollaston T. V. 1864: Catalogue of the coleopterous insects of the Canaries in the collection of the British Museum. London: Trustees of the British Museum, xiii + 648 pp.
Wollaston T. V. 1865: Coleoptera Atlantidum, being an enumeration of the coleopterous insects of the Madeiras, Salvages and Canaries. London: John van Voorst, xlvii + 526 pp + appendix 140 pp.
SELECTED ENTOMOLOGICAL LINKS
LINKS WITHIN THIS WEBSITE
→ Scientific publications (pdf)
→ Colleagues’ Gallery
→ Joy of Entomology
→ Macrophotography
→ Insects at Scale
→ Anatomy of a Weevil
INSTITUTIONS
→ NHM (London)
→ MNCN (Madrid)
→ MCN (Tenerife)
→ Curculio Institut
→ Spanish Entomology Association
→ Melansis – SEC