Beetles, beetles, beetles …

Lizards, snails, caterpillars and beetles were common toys to play with when I was a child and enjoyed the countryside in La Palma or at the farm in Tenerife. Later, when I was a teenager I started a collection of beetles, and since then, I haven´t stopped, Entomology having become a Leitmotiv and umbilical cord with nature up to the present. Beetles were probably the hidden reason why I decided to study Biology at the University; when the pressure at home –a mild one– was to do something more “serious”, like Architecture, Agrarian engineering or Philosophy.  However, I have never worked in Entomology professionally; but always as a happy and priviledged amateur.


Entomological cabinet of Dr. A. Machado

When I finished my Biology Degree in La Laguna, the knowledge about Entomology of the Canary Islands was dispersed mainly in foreign institutions and journals. In 1976 I established, along with my wife and my brother, a cultural non-profit private association IBCER (International Bureau of Canarian Entomological Research) with the aim of collecting that information and offering a contact point for all researchers interested in our insect fauna.

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IBCER was more or less useful until the Department of Zoology at the University of La Laguna, and the Natural History Museum of Santa Cruz consolidated their activity in Entomology during the 80,s. In January 1994, IBCER had 53 colleagues registered.

At present, IBCER is dormant, but my library stays open and the same spirit of assistance to any colleague persists in the second floor at home, where I have established my cabinet (you can see a set of photos at the end of this page). Now, I have the intention to provide as much information as I can through this web-site, mainly on Coleoptera, bibliography, and whatever is in my hands, with the only limit being that of available time.

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Studying the Wollaston Collection

When you are young and looking at bugs, each group has attractive aspects: ants, springtails, spiders, grasshoppers, dragonflies… My interest in ground-beetles goes back to 1969, when I discovered a few black Microlestes close to my golf ball. Its perfect design just fascinated me. The choice was made and it received full support of Don José María Fernández, curator of the Museum of Natural History at Santa Cruz, whom I started to visit regularly.

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Later, in 1975, I presented my Biology Diploma work on the ground-beetles of the laurel forest, and that was the start of a more ambitious project in which I intended to answer questions like: How many species do really live in the Canary Islands? Are they all valid species? Where do they live? It took me about 15 years to perform a full taxonomic and faunal revision to aarive at some satisfactory answers. That was my PhD dissertation (cum laude).

I studied some 23.000 specimens from my own collection as well as from several museum collections (Madrid, Oxford, London, Paris, etc.). A total of 232 species live in the Canary Islands, 70% of which are endemic to the archipelago. I revised all the endemic taxa and described 1 tribe, 4 new genera and 29 new species. In parallel, I established 28 new synonyms. The Monografía de los carábidos de las islas Canarias, was published a year later (1992). It has 734 pages and 313 figures. Some chapters deal with aspects like phenology, dispersal power, and parasites, with some emphasis on biogeography and comments on evolution. But at that time, I did not perform phylogenetic analysis. That came later, but not with carabids.

You can access the Monograph here (PDF) or from the section Publication /Scientific, were all other papers dealing with Carabidae are available.

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Having grown after the monograph on Carabidae, my interest in understanding how species form and why, islands are the best places to study evolution, and beetles the best candidates. I defined a profile for the ideal group: it has to be monophyletic, with as many species as possible, all endemic and without flying capacity. Winner by a wide margin: the genus Laparocerus Schönherr, 1834 (Curculionidae: Entiminae).

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In 1999 I switched from ground-beetles to weevils and designed a project to search for Laparocerus in all of Macaronesia, review the taxonomy, study their ecology and reveal their phylogeny and paths of evolution. If there are any patterns, they should be expressed there.

After 15 years of studying the genus I am still not finished, but getting closer to the end. The Laparocerus are not as rare as previously thought. They are easy to be found by night. I developed efficient techniques for sampling, having collected more than 32.000 specimens, many of them belonging to species previously unknown to science. I prospected the Azores, Madeira and Canary Islands consciously, repeating expeditions to the same island, or to the same place whenever needed. Additionally, I have studied the material kept in all major collections in European museums, plus that sent by colleagues. If you have a recalcitrant species of Laparocerus to be identified, please do not hesitate to write to me.

In the section of Production/Scientific there are several articles about Laparocerus, many describing new species. The number of species level taxa has increased from 123 (when I started) to 215, and there are some more in the pipeline.

I faced the phylogenetic study with the collaboration of Dr. Mariano Hernández Ferrer, from the University Institute of Tropical Diseases and Public Health of the Canary Islands, and hired some technical assistance for laboratory work. For financing part of the costs of the molecular study I received a subvention in 2004-2005 from the Fundación Biodiversidad (Madrid). Needless to say, I had to update my knowledge of Genetics and studied the new methodologies for phylogenetic inference and a set of complex software, all with the good guidance of Mariano. I learned a lot –what I always appreciate– , and I am convinced that he also learned something about beetles. The phylogeny of the Madeiran clade was published in 2008, and that of the Canarian-Moroccan clade should be finished in 2015.

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Catalogue of Canarian Coleoptera

Wollaston Collection

Since Coleoptera Atlantidum of Wollaston (1865), by the end of the XX Century, there was no modern account of beetles from the Canary Islands. In collaboration with Dr. Pedro Oromí, from the University of La Laguna, and with the assistance of many colleagues, we published a new catalogue in 2000 (see PDF), updating to 2061 the total number of species and subspecies, of which 1247 are endemic. We also registered 325 recognized synonyms.

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I keep the Coleoptera database alive, and Dr. Oromí is in charge of Entomology in the Banco de datos de Biodiversidad de Canarias (BIOTA), supported by the Canarian Government. Should you have any amendment or contribution to add to the Canarian Coleoptera Database, please feel welcome to contact one of us:

When I find a suitable database and some time, it is my intention to include the Canarian Catalogue in this website for online access. For the moment, I keep it in my desktop running on Procite, and oldie, but performing well.

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Coleoptera Database (coming soon)

Canarian Entomological Bibliography

Books on Coleoptera by T. Vernon Wollaston

One of my main goals at IBCER was to find all the entomological bibliography related to the Canary Islands. That was in the times of cards, typing machines, carbon copies, ordinary letters and smailmail to contact other colleagues, visiting libraries in person, etcetera; something unimaginable for the Internet generation. In 1976 I finished a preliminary list, and in 1987 the book A Bibliography of the Canary Island Entomology was published, which was a milestone.

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Since then, I have been registering all new publications as best as I can. They have grown exponentially, but the new facilities with computers and Internet is a great help. As with the Catalogue of Coleoptera, I hope to be able to transfer my database to an online application within this web-site (probably RefBASE). It is a question of finding the time. Meanwhile, if you need information about any publication, please write to me. In some cases, I can also provide a PDF or scan the original. My library has some 12.000 registers (EndNote X6), and about 4800 are related with Arthropods.

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Bibliography Database (coming soon)

How to get a Collecting Permit for Insects

Looking for beetles below stones

Collecting insects or any other biological material has become complicated in the Canary Islands. The Spanish national legislation on wildlife conservation and in particular the authorization process for collecting samples was designed with large mammals or birds in the mind, not with insects. To get a permit is possible, but it takes time. Many colleagues just write to me, but I do not extend permits. What I can do is help with the bureaucracy and cross fingers.

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Almost half of the territory of the Canary Islands is covered by some kind of protected area (natural monuments, national parks, rural parks, strict reserves, etc.). To collect in any of these areas, you will need a permit from the conservation authority, which is at present the Island Council (Cabildo Insular) of the respective island, except for national parks (Park Authority). Yes, you will need a permit for each island or national park, if you intend to work on more than one.

Below you will find a link to a new page with application forms, contacts, and some recommendations. Not all Cabildos will ask for the same information, but it refers basically to questions like: what, where, when, how, who, and what for. It will take X time to get a reply, and it is common that the authorization includes some conditions.

If some of your target species are protected because they are threatened, the application procedures are different and the Biodiversity Service of the Autonomous Government will be involved.

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Application forms & guidelines







Director: Antonio Machado Carrillo
Filmed on: Super 8, original footage
© 1971: Antonio Machado Carrillo


Director: Antonio Machado Carrillo
Location: Tenerife, Canary Islands
© 2020: Antonio Machado Carrillo


Director: Antonio Machado Carrillo
Location: Tenerife, Canary Islands
© 2020: Antonio Machado Carrillo




  1. Dicrodontini Mach. 1992, Carabidae: Zuphiinae


  1. Canarobius Mach. 1987, Carabidae, Trechinae, Trechodini
  2. Spelaeovulcania Mach. 1987, Carabidae, Trechinae, Trechodini
  3. Wolltinerfia Mach. 1985 nom.nov. pro Wollastonia Mach. 1984, Carabidae Pterostichinae
  4. Orzolina Mach. 1987c, Carabidae, Trechinae, Bembidini


  1. Gietopus 1992, del género Wolltinerfia Mach. 1985, Carabidae
  2. Lauricalathus 1992, del género Calathus Bon., Carabidae
  3. Fortunotrox  2011, del género Laparocerus Schoenh. 1834, Curculionidae
  4. Pseudatlantis  2008, del género Laparocerus Schoenh. 1834, Curculionidae
  5. Wollastonius 2008, del género Laparocerus Schoenh. 1834, Curculionidae
  6. Atlantodes  2008, del género Laparocerus Schoenh. 1834, Curculionidae
  7. Faycanius Mach. 2012, del género Laparocerus Schoenh. 1834, Curculionidae
  8. Aridotrox , 2017, del género Laparocerus Schoenh. 1834, Curculionidae
  9. Belicarius , 2017, del género Laparocerus Schoenh. 1834, Curculionidae
  10. Bencomius, 2017, del género Laparocerus Schoenh. 1834, Curculionidae
  11. Canariotrox , 2017, del género Laparocerus Schoenh. 1834, Curculionidae
  12. Purpuranius, 2017, del género Laparocerus Schoenh. 1834, Curculionidae


  1. Pimelia fernandezlopezi Mach. 1979, Tenebrionidae.
  2. Canarobius chusyae Mach. 1988. Carabidae
  3. Canarobius oromii Mach. 1988. Carabidae
  4. Thalassophilus subterraneus Mach. 1989. Carabidae
  5. Spelaeovulcania canariensis Mach. 1987. Carabidae
  6. Trechus minioculatus Mach. 1987. Carabidae
  7. Trechus benahoaritus Mach. 1989. Carabidae
  8. Lymnastis subovatus Mach. 1992. Carabidae
  9. Lymnastis thoracicus Mach. 1992. Carabidae
  10. Tachys fortunatus Mach. 1989. Carabidae
  11. Orzolina thalassophila Mach. 1987. Carabidae
  12. Wolltinerfia tenerifae (Mach. 1984). Carabidae (sub. Wollastonia)
  13. Wolltinerfia martini (Mach. 1984). Carabidae (sub. Eutrichopus)
  14. Calathus refleximargo Mach. 1992. Carabidae
  15. Calathus pilosipennis Mach. 1992. Carabidae
  16. Calathidius brevithorax Mach. 1992. Carabidae
  17. Licinopsis angustula Mach. 1987. Carabidae
  18. Licinopsis schurmanni Mach. 1987. Carabidae (como ssp. de L. angustula)
  19. Zabrus canaricus Mach. 1992. Carabidae
  20. Bradycellus feloi Mach. 1992. Carabidae [= syn.]
  21. Paradromius iucundus Mach. 1992. Carabidae
  22. Paradromius tamaranus Mach. 1992. Carabidae
  23. Paradromius amplius Mach. 1992. Carabidae
  24. Paradromius exornatus Mach. 1992. Carabidae
  25. Paradromius purpurarius Mach. 1992. Carabidae
  26. Paradromius hariensis Mach. 1992. Carabidae
  27. Paradromius scholzi Mach. 1992. Carabidae
  28. Philorhizus mateui Mach. 1992. Carabidae
  29. Philorhizus franzi Mach. 1992. Carabidae
  30. Metadromius palmi Mach. 1992. Carabidae
  31. Parazuphium feloi Mach. 1998. Carabidae
  32. Drouetius borgesi Mach. 2009. Curculionidae
  33. Drouetius oceanicus Mach. 2009. Curculionidae
  34. Laparocerus bacalladoi Mach. 2005. Curculionidae
  35. Laparocerus junonius Mach., 2007. Curculionidae
  36. Laparocerus aguiari Mach. 2007. Curculionidae
  37. Laparocerus subparalellus Mach., 2007. Curculionidae
  38. Laparocerus boticarius Mach., 2007. Curculionidae
  39. Laparocerus roudieri Mach., 2007. Curculionidae
  40. Laparocerus chasnensis Mach.,2007. Curculionidae
  41. Laparocerus tinguaro Mach., 2007. Curculionidae
  42. Laparocerus humeralis Mach., 2007. Curculionidae
  43. Laparocerus acutipennis Mach., 2007. Curculionidae
  44. Laparocerus exiguus Mach., 2007. Curculionidae
  45. Laparocerus anopthalmus Mach., 2007. Curculionidae
  46. Laparocerus hupalupa Mach., 2007. Curculionidae
  47. Laparocerus orone Mach., 2007. Curculionidae
  48. Laparocerus oculatissimmus Mach., 2007. Curculionidae
  49. Laparocerus depressus Mach., 2007. Curculionidae
  50. Laparocerus spinimanus Mach., 2007. Curculionidae
  51. Laparocerus mulagua Mach., 2007. Curculionidae
  52. Laparocerus benchijigua Mach., 2007. Curculionidae
  53. Laparocerus inermis Mach., 2007. Curculionidae
  54. Laparocerus heres Mach., 2007. Curculionidae
  55. Laparocerus madeirensis, Mach. 2008. Curculionidae
  56. Laparocerus edaphicus Mach. 2008. Curculionidae
  57. Laparocerus prainha Mach. 2008. Curculionidae
  58. Laparocerus oromii Mach. 2008. Curculionidae
  59. Laparocerus stuebeni Mach. 2008. Curculionidae
  60. Laparoceerus hobbit Mach. 2008. Curculionidae
  61. Laparocerus cryptus Mach. 2008. Curculionidae
  62. Laparocerus silvaticus Mach. 2008. Curculionidae
  63. Laparocerus serrado Mach. 2008. Curculionidae
  64. Laparocerus lopezi Mach. 2008. Curculionidae
  65. Laparocerus aeneotincutus Mach. 2009. Curculionidae
  66. Laparocerus elongatus Mach. 2009. Curculionidae
  67. Laparocerus supranubius Mach. 2009. Curculionidae
  68. Laparocerus astralis Mach. 2009. Curculionidae
  69. Laparocerus amicorum Mach. 2009. Curculionidae
  70. Laparocerus cristatus Mach. 2009. Curculionidae
  71. Laparocerus feloi Mach. 2009. Curculionidae
  72. Laparocerus morrisi Mach. 2009. Curculionidae
  73. Laparocerus decipiens Mach. 2009. Curculionidae
  74. Laparocerus tanausu Mach. 2009. Curculionidae
  75. Laparocerus arrochai Mach. 2009. Curculionidae
  76. Laparocerus acyphus Mach. 2009. Curculionidae
  77. Laparocerus femoralis Mach. 2009. Curculionidae
  78. Laparocerus mucronatus Mach. 2009. Curculionidae
  79. Laparocerus tarsalis Mach. 2009. Curculionidae
  80. Laparocerus cephalotes Mach. 2011. Curculionidae
  81. Laparocerus bimbache Mach. 2011. Curculionidae
  82. Laparocerus cavernarius Mach. 2011. Curculionidae
  83. Laparocerus hypogeus Mach. 2011. Curculionidae
  84. Laparocerus rotundatus Mach. 2011. Curculionidae
  85. Laparocerus pilosiventris Mach. 2011. Curculionidae
  86. Laparocerus confusus Mach. 2011. Curculionidae
  87. Laparocerus juelensis Mach. 2011. Curculionidae
  88. Laparocerus magnificus Mach. 2011. Curculionidae
  89. Laparocerus iruene Mach. & García 2011. Curculionidae
  90. Tarphius jandiensis Mach. 2011. Zopheridae
  91. Tarphius desertaensis Mach. 2011. Zopheridae
  92. Laparocerus maxorata Mach. 2011. Curculionidae
  93. Laparocerus colonnellii Mach. 2011. Curculionidae
  94. Laparocerus calvus Mach. 2011. Curculionidae
  95. Laparocerus xericola Mach. 2011. Curculionidae
  96. Laparocerus fraterculus Mach. 2011. Curculionidae
  97. Laparocerus longipennis Mach. 2011. Curculionidae
  98. Laparocerus estevezi Mach. 2012. Curculionidae
  99. Laparocerus arcanus Mach. 2012. Curculionidae
  100. Laparocerus dissidens Mach. 2012. Curculionidae
  101. Laparocerus bentejui Mach. 2012. Curculionidae
  102. Laparocerus anniversarius Mach. 2012. Curculionidae
  103. Laparocerus franzi Mach. 2012. Curculionidae
  104. Laparocerus teldense Mach. 2012. Curculionidae
  105. Laparocerus fraudulentus Mach. 2012. Curculionidae
  106. Laparocerus hytricoides Mach. 2012. Curculionidae
  107. Laparocerus rugosivertex Mach. 2012. Curculionidae
  108. Laparocerus ornatus Mach. 2012. Curculionidae
  109. Laparocerus marmoratus Mach. 2012. Curculionidae
  110. Laparocerus tirmensis Mach. 2012. Curculionidae
  111. Laparocerus semipilosus Mach. 2012. Curculionidae
  112. Laparocerus osorio Mach. 2012. Curculionidae
  113. Laparocerus tirajana Mach. 2012. Curculionidae
  114. Oromia thoracica Mach. & López, 2015. Curculionidae
  115. Laparocerus merigensis Mach. 2015. Curculionidae
  116. Laparocerus dilutus Mach., 2015. Curculionidae
  117. Laparocerus notatus Mach., 2015. Curculionidae
  118. Laparocerus teselinde Mach., 2015. Curculionidae
  119. Laparocerus quadratus Mach., 2015. Curculionidae
  120. Laparocerus macilentus Mach., 2015. Curculionidae
  121. Laparocerus sonchiphagus Mach., 2015. Curculionidae
  122. Laparocerus tenicola Mach., 2015. Curculionidae
  123. Laparocerus anagae Mach., 2015. Curculionidae
  124. Laparocerus campestris Mach., 2015. Curculionidae
  125. Laparocerus abona Mach., 2016. Curculionidae
  126. Laparocerus tauce Mach., 2016. Curculionidae
  127. Laparocerus crotchi Mach., 2016. Curculionidae
  128. Laparocerus gerodes Mach, 2016. Curculionidae
  129. Laparocerus tafadensis Mach., 2016. Curculionidae
  130. Laparocerus soniae Mach., 2016. Curculionidae
  131. Laparocerus auarita Mach., 2016. Curculionidae
  132. Laparocerus canescens Mach., 2016. Curculionidae
  133. Laparocerus punctiger Mach., 2016. Curculionidae
  134. Laparocerus sanctaecrucis Mach,, 2016. Curculionidae
  135. Laparocerus federico Mach. 2018. Curculionidae
  136. Herpisticus famarae Mach. 2020. Curculionidae
  137. Herpisticus betancuriae Mach. 2020. Curculionidae
  138. Herpisticus jandiensis Mach. 2020. Curculionidae
  139. Herpisticus rectipes Mach. 2020. Curculionidae
  140. Herpisticus guanarteme Mach. 2020. Curculionidae
  141. Herpisticus denudatus Mach. 2020. Curculionidae
  142. Herpisticus gigas Mach. 2020. Curculionidae
  143. Herpisticus guayarmina Mach. 2020. Curculionidae
  144. Herpisticus tasarticus Mach. 2020. Curculionidae
  145. Herpisticus scopulus Mach. 2020. Curculionidae
  146. Herpisticus nanus Mach. 2020. Curculionidae
  147. Herpisticus daute Mach. 2020. Curculionidae
  148. Herpisticus aridicola Mach. 2020. Curculionidae
  149. Herpisticus gomerensis Mach. 2020. Curculionidae
  150. Herpisticus hispidus Mach. 2020. Curculionidae
  151. Herpisticus bobadillae Mach. 2020. Curculionidae


  1. Aepus gracilicornis meridionalis Mach. 1992. Carabidae
  2. Olisthopus glabratus tamaranus Mach. 1992. Carabidae
  3. Platyderus alticola hierroensis Mach. 1992. Carabidae
  4. Platyderys alticola gomeraensis Mach. 1992. Carabidae
  5. Licinopsis obliterata franzi Mach. 1987. Carabidae
  6. Licinopsis picescens gomerita Mach. 1987. Carabidae
  7. Paradromius iucundus gillerforsi Mach. 1992. Carabidae
  8. Paradromius exornatus furvus Mach. 1992. Carabidae
  9. Paradromius strigifrons grancanariensis Mach. 1992. Carabidae
  10. Paradromius insularis illautus Mach. 1992. Carabidae
  11. Dromius angustus dissimilis Mach. 1992. Carabidae
  12. Philorhizus atlanticus palmensis Mach. 1992. Carabidae
  13. Philorhizus incertus jandiensis Mach. 1992. Carabidae
  14. Dicrodontus brunneus exilis Mach. 1992. Carabidae
  15. Parazuphium damascenum canariense Mach. 1992. Carabidae
  16. Drouetius azoricus nitens Mach. 2009. Curculionidae
  17. Drouetius azoricus parallelirostris Mach. 2009. Curculionidae
  18. Drouetius azoricus separandus Mach. 2009. Curculionidae
  19. Drouetius borgesi centralis Mach. 2009. Curculionidae
  20. Drouetius borgesi sanctimichaelis Mach. 2009. Curculionidae
  21. Drouetius oceanicus tristis Mach. 2009. Curculionidae
  22. Laparocerus aethiops garajonay Mach., 2007. Curculionidae
  23. Laparocerus heres jocoensis Mach., 2007. Curculionidae
  24. Laparocerus mateui tuberosus Mach. 2011. Curculionidae
  25. Laparocerus orone aridane Mach. 2009. Curculionidae
  26. Laparocerus orone hierroensis Mach. 2011. Curculionidae
  27. Laparocerus zarazagai subreflexus Mach. & García 2011. Curculionidae
  28. Laparocerus rasus jandiensis Mach. 2011. Curculionidae
  29. Laparocerus rasus betancor Mach. 2011. Curculionidae
  30. Laparocerus curvipes famara Mach. 2011. Curculionidae
  31. Laparocerus garretai albosquamosus Mach. 2011. Curculionidae
  32. Laparocerus susicus inexpectatus Mach. 2011. Curculionidae
  33. Laparocerus susicus montanus Mach. 2011. Curculionidae
  34. Laparocerus squamosus tasarticus Mach. 2012. Curculionidae
  35. Laparocerus alluaudi aytamis Mach. 2012. Curculionidae
  36. Laparocerus bentejui delicatulus Mach. 2012. Curculionidae
  37. Laparocerus bentejui robustus Mach. 2012. Curculionidae
  38. Laparocerus subnodosus montivagans Mach. 2013. Curculionidae
  39. Laparocerus sanchezi arures , 2016. Curculionidae
  40. Laparocerus obscurus daute , 2016. Curculionidae
  41. Laparocerus tinguaro tabornoi , 2016. Curculionidae
  42. Laparocerus dissimilis alticola , 2016. Curculionidae
  43. Laparocerus dissimilis infernalis , 2016. Curculionidae
  44. Laparocerus tenuepunctatus oppositus , 2016. Curculionidae
  45. Herpisticus subvestitus pseudolanatus Mach. 2020. Curculionidae
  46. Herpisticus hierrensis benahoare Mach. 2020. Curculionidae
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Subfamily Entiminae Schoenherr, 1823

Tribe Laparocerini Lacordaire, 1863

Genus Laparocerus Schoenherr, 1834: 530 type species Laparocerus morio Boheman, 1834

              = Cyphoscelis Wollaston, 1854: 356 type species Cyphoscelis distorta Wollaston, 1854

Subgenus Amyntas Wollaston, 1865: Appendix, 55  type species Lichenophagus incomptus Wollaston, 1865

Canopus Wollaston, 1864: 337 [HN] type species Atlantis subnebulosa Wollaston, 1864

arrochai Machado, 2009: 190 N: CI (La Palma)

bellus Roudier, 1957a: 31 N: CI (Tenerife)

cristatus Machado, 2009: 193 N: CI (La Palma)

fernandezi Roudier, 1957a: 34 N: CI (Tenerife)

incomptus Wollaston, 1865: Appendix, 55 (Lichenophagus) N: CI (El Hierro)

ruteri Roudier, 1957a: 33 N: CI (Tenerife)

subnebulosus Wollaston, 1864: 337 (Atlantis) N: CI (Gran Canaria)

tanausu Machado, 2009: 195 N: CI (La Palma)

tetricus Boheman, 1834: 542 (Eremnus) N: CI (Tenerife)

simplex Brullé, 1839: 71 (Otiorhynchus)

tibialis Wollaston, 1864: 338 (Atlantis) N: CI (Tenerife)

uyttenboogaarti Zumpt, 1940: 677 N: CI (Tenerife)

Subgenus Anillobius Fauvel, 1907: 71 type species Anillobius solifuga Fauvel, 1907

solifuga Fauvel, 1907: 72 (Anillobius) N: MR (Madeira)

portosantoi Franz 1970: 3 (Anillobius) N: MR (Porto Santo)

Subgenus Aridotrox Machado 2017: 33 type species Laparocerus rasus Wollaston, 1864

colonnellii Machado, 2011b: 232 N: CI (Fuerteventura)

dispar Wollaston, 1864: 357 N: CI (Lanzarote)

rasus betancor Machado, 2011b: 223 N: CI (Fuerteventura)

rasus jandiensis Machado, 2011b: 224 N: CI (Fuerteventura)

rasus rasus Wollaston, 1864: 354 N: CI (Lanzarote)

susicus inexpectatus Machado, 2011b: 229 N: MO

tessellatus Hustache, 1932: 43 (Cyclomaurus) [HN]

susicus montanus Machado, 2011b: 229 N: MO

susicus susicus Escalera, 1914: 424 (Cyclobarus) N: MO

xericola Machado, 2011b: 231 N: CI (Fuerteventura)

Subgenus Atlantis Wollaston, 1854: 361 type Atlantis lamellipes Wollaston, 1854

calcatrix Wollaston, 1854: 366 (Atlantis) N: MR (Madeira)

calcator Marseul, 1872: 546

clavatus Wollaston, 1854: 363 (Atlantis) N: MR (Madeira)

hobbit Machado, 2008a: 318 N: MR (Madeira)

lamellipes Wollaston, 1854: 364 (Atlantis) N: MR (Madeira)

lauripotens Wollaston, 1854: 369 (Atlantis) N: MR (Madeira)

              = australis Wollaston, 1864: 370 (Atlantis)

lindbergi Roudier, 1963: 133 N: MR (Madeira)

madeirensis Machado, 2008a: 316 N: MR (Madeira)

noctivagans Wollaston, 1854: 367 (Atlantis) [NP] N: MR (Madeira)

                  = piceus Boheman, 1834: 531 [NO]

serrado Machado, 2008a: 319 N: MR (Madeira)

undulatus Wollaston, 1862: 289 N: MR (Madeira)

vespertinus Wollaston, 1854: 371 (Atlantis) N: MR (Madeira)

Subgenus Atlantodes Machado, 2008a: 312 type species Atlantis mendax Wollaston, 1854

colasi Roudier, 1958: 205 N: MR (Madeira)

inconstans Wollaston, 1854: 375 (Atlantis) N: MR (Porto Santo)

instabilis Wollaston, 1854: 377 (Atlantis) N: MR (Porto Santo)

lanatus Wollaston, 1854: 372 (Atlantis) N: MR (Madeira)

mendax Wollaston, 1854: 376 (Atlantis) N: MR (Porto Santo)

navicularis Wollaston, 1854: 374 (Atlantis) N: MR (Porto Santo)

prainha Machado, 2008a: 323 N: MR (Madeira)

Subgenus Belicarius Machado 2017: 35 type species Laparocerus mendicus Wollaston, 1864

bentejui bentejui Machado, 2012: 7 N: CI (Gran Canaria)

bentejui delicatulus Machado, 2012: 8 N: CI (Gran Canaria)

bentejui robustus Machado, 2012: 10 N: CI (Gran Canaria)

exiguus Machado, 2007b: 11 N: CI (La Gomera)

exophthalmus Machado, 2007b: 12 N: CI (La Gomera)

feloi Machado, 2009: 203 N: CI (La Palma)

longiclava Lindberg, 1953: 15 N: CI (Tenerife)

magnificus Machado, 2011a: 70 N: CI (La Gomera)

mateui mateui Roudier, 1954: 85 N: CI (La Gomera)

mateui tuberosus Machado, 2011a: 74 N: CI (El Hierro)

mendicus Wollaston, 1864: 355 N: CI (El Hierro)

morrisi Machado, 2009: 218 N: CI (La Palma)

oculatissimus Machado, 2007b: 14 N: CI (La Gomera)

rotundatus Machado, 2011a: 65 N: CI (La Gomera)

sanchezi arures Machado, 2016: 33 N: CI (La Gomera)

sanchezi sanchezi Roudier, 1957a: 24 N: CI (El Hierro)  

              = nitidirostris Roudier, 1957a: 25

Subgenus Bencomius Machado 2017: 35 type species Laparocerus grossepunctatus Wollaston, 1864

bolivari Uyttenboogaart, 1937: 97 N: CI (Tenerife)

boticarius Machado, 2007a: 39 N: CI (Tenerife)

combrecitensis Roudier, 1957c: 90 [RNJ N: Cl (La Palma)

palmensis Roudier, 1957a: 20 [HN]

crassifrons Wollaston, 1864: 345 N: CI (Tenerife)

edaphicus Machado, 2008b: 1278 N: CI (Tenerife)

escaleraorum Uyttenboogaart, 1937: 96 N: CI (Tenerife)

gomerensis Lindberg, 1953: 15 N: CI (La Gomera)

grossepunctatus Wollaston, 1864: 344 N: CI (Tenerife)

scapularis Wollaston, 1864: 347 N: CI (Tenerife)

wollastoni Marseul, 1872: 521

subparallelus Machado, 2007a: 43 N: CI (Tenerife)

supranubius Machado, 2009: 210 N: CI (La Palma)

tenuepunctatus oppositus Machado, 2016: 16 N: CI (Tenerife)

tenuepunctatus tenuepunctatus Roudier, 1957a: 20 N: CI (Tenerife)

undatus Wollaston, 1864: 342 N: CI (Tenerife)

Subgenus Canariotrox Machado 2017: 35 type species Laparocerus inaequalis Wollaston, 1864

abona Machado, 2016: 14 N: CI (Tenerife)

acyphus Machado, 2009: 189 N: CI (La Palma)

aeneotinctus Machado, 2009: 185 N: CI (La Palma)

affinis Wollaston, 1864: 358 N: CI

aguiari Machado, 2007a: 36 N: CI (Tenerife)

crassus Roudier, 1957a: 28 N: CI (Tenerife)

estevezi Machado, 2012: 13N: CI (Gran Canaria)

femoralis Machado, 2009: 188 N: CI (La Palma)

hirtus Wollaston, 1864: 348 N: CI (Gran Canaria)

              = doramasensis Uyttenboogart, 1929: 352

inaequalis globulipennis Wollaston, 1864: 349 N: CI (La Palma)

globosipennis Marseul, 1872: 531

inaequalis inaequalis Wollaston, 1863: 220 N: CI (Tenerife)

occidentalis Wollaston, 1864: 350 N: CI (El Hierro)

rugosicollis Uyttenboogaart, 1937: 98 N: CI (Tenerife)

tauce Machado, 2016: 16 N: CI (Tenerife)

vestitus Wollaston 1864: 358 N: CI

velatus Marseul, 1872: 532

Subgenus Faycanius Machado, 2012: 29 type species Otiorhynchus squamosus Brullé, 1839

alluaudi alluaudi Uyttenboogaart, 1940: 55 N: CI (Gran Canaria)

alluaudi aytamis Machado, 2012: 33 N: CI (Gran Canaria)

anniversarius Machado, 2012: 37 N: CI (Gran Canaria)

arcanus Machado, 2012: 32 N: CI (Gran Canaria)

dissidens Machado, 2012: 35 N: CI (Gran Canaria)

squamosus squamosus Brullé, 1839: 71 (Otiorhynchus) N: CI (Gran Canaria)

seniculus Wollaston, 1864: 353

squamosus tasarticus Machado, 2012: 21 N: CI (Gran Canaria)

Subgenus Fernandezius Roudier, 1957a: 40 type species Lichenophagus tesserula Wollaston, 1864

anagae Machado, 2015: 126 N: CI (Tenerife)

campestris Machado, 2015: 132 N: CI (Tenerife)

impressicollis Wollaston, 1864: 368 (Lichenophagus) N: CI (Tenerife)

macilentus Machado, 2015: 125 N: CI (Tenerife)

persimilis Wollaston, 1864: 365 (Lichenophagus) N: CI (Tenerife)

sculptipennis montivagans Machado 2013: 69  N: CI (La Palma)

              = palmensis Roudier 1957a: 43 (Lichenophagus) [HN]

sculptipennis sculptipennis Wollaston, 1864: 367 (Lichenophagus) N: CI (La Palma)

seriesetosus Wollaston, 1864: 365 (Lichenophagus) N: CI [La Palma]

sonchiphagus Machado, 2015: 120 N: CI (Tenerife)

subcalvus Wollaston, 1864: 336 (Lichenophagus) N: CI (El Hierro)

subnodosus Wollaston, 1864: 336 (Lichenophagus) N: CI (Tenerife)

tenicola Machado, 2015: 128 N: CI (Tenerife)

tesserula Wollaston, 1864: 364 (Lichenophagus) N: CI (Tenerife)

Subgenus Fortunotrox Machado, 2011a: 58 type species Laparocerus gracilis Wollaston, 1865

acutipennis Machado, 2007b: 5 N: CI (La Gomera)

benchijigua Machado, 2007b: 6 N: CI (La Gomera)

confusus Machado, 2011a: 62 N: CI (La Gomera)

crotchi Machado, 2016: 30 N: CI (La Gomera)

decipiens Machado, 2009: 214 N: CI (La Palma)

depressus Machado, 2007b: 7 N: CI (La Gomera)

gracilis Wollaston, 1864: 356 N: CI (La Gomera)

gomeraecola Marseul, 1872: 531

indutus Wollaston, 1865 (app.): 53 N: CI (La Gomera)

tectus Marseul, 1872: 535

juelensis Machado, 2011a: 64 N: CI (La Gomera)

orone aridane Machado, 2009: 211 N: CI (La Palma)

orone hierrensis Machado, 2011a: 80 N: CI (El Hierro)

orone orone Machado, 2007b: 3 N: CI (La Gomera)

puncticollis Wollaston, 1864: 362 N: CI (El Hierro)

spinimanus Machado, 2007b: 9 N: CI (La Gomera)

subopacus Wollaston, 1865 (app.): 52 N: CI (La Gomera)

Subgenus Guanchotrox Alonso-Zarazaga & Lyal, 1999: 11 [RN] type species Laparocerus canariensis Boheman, 1843

              = Amphora Wollaston, 1864: 335 [HN] type species Laparocerus canariensis Boheman, 1843

amicorum Machado, 2009: 212 N: CI (La Palma)

astralis Machado, 2009: 206 N: CI (La Palma)

buenavistae Roudier, 1957a: 29 N: CI (Tenerife)

canariensis Boheman, 1842: 228 N: CI (Tenerife)

cephalotes Machado, 2011a: 78 N: CI (El Hierro)

debilis Wollaston, 1865: Appendix, 53 N: CI (Tenerife)

dissimilis alticola Machado, 2016: 22 N: CI (Tenerife)

dissimilis infernalis Machado 2016: 23 N: CI (Tenerife)

dissimilis dissimilis Lindberg, 1950: 17 N: CI (Tenerife)

elongatus Machado, 2009: 198 N: CI (La Palma)

gerodes Machado, 2016: 23 N: CI (La Gomera)

humeralis Machado, 2007b: 23 N: CI (La Gomera)

junonius Machado, 2007b: 19 N: CI (La Gomera)

marmoratus Machado, 2012: 22 N: CI (Gran Canaria)

mucronatus Machado, 2009: 201 N: CI (La Palma)

obscurus daute Machado, 2016: 21 N: CI (Tenerife)

obscurus obscurus Wollaston, 1864: 355 N: CI (Tenerife)

              = obscurellus Marseul, 1872: 539

obtriangularis Wollaston, 1864: 351 N: CI (Tenerife)

ornatus Machado, 2012:20 N: CI (Gran Canaria)

palmensis Lindberg, 1953: 16 N: CI (La Palma)

seminitens Lindberg, 1950: 17 N: CI (Tenerife)

tafadensis Machado, 2016: 28 N: CI (Tenerife)

tarsalis Machado, 2009: 205 N: CI (La Palma)

tenellus Wollaston, 1864: 362 N: CI (Tenerife)

tirmensis Machado, 2012: 21 N: CI (Gran Canaria)

roudieri Machado, 2007b: 22 N: CI (La Gomera)

tinguaro tabornoi Machado, 2016: 29 N: CI (Tenerife)

tinguaro tinguaro Machado, 2007a: 44 N: CI (Tenerife)

transversus Lindberg, 1950: 16 N: CI (Tenerife)

Subgenus Laparocerus Schoenherr, 1834: 530 type species Laparocerus morio Boheman, 1834

              = Cyphoscelis Wollaston, 1854: 356 type species Cyphoscelis distorta Wollaston, 1854

chaoensis cevadae Roudier, 1961: 62 N: MR (Ilheu do Desembarcadouro)

chaoensis chaoensis Uyttengoogaart, 1940: 57 N: MR (Ilheu de Chão)

              = vandeli Roudier 1958: 199

cryptus Machado, 2008a: 315 N: MR

distortus Wollaston, 1854: 357 (Cyphoscelis) N: MR (Madeira)

morio Boheman, 1834: 531 N: MR (Madeira)

obesulus Desbrochers, 1904: 63

Subgenus Lichenophagus Wollaston, 1854: 389 type species Lichenophagus fritillus Wollaston, 1854

acuminatus Wollaston, 1854: 391 E: MR (Porto Santo)

fritillus Wollaston, 1854: 390 E: MR (Deserta Grande)

Subgenus Machadotrox Alonso-Zarazaga & Lyal, 1999: 10 type species Otiorhynchus sculptus Brullé, 1839

aethiops aethiops Wollaston, 1864: 347 N: CI (El Hierro)

aethiops garajonay Machado, 2007b: 15 N: CI (La Gomera)

cavernarius Machado, 2011a: 84 N: CI (El Hierro)

chasnensis Machado, 2007a: 41 N: CI (Tenerife)

dacilae García, 1988: 45 N: CI (La Palma)

excavatus excavatus Wollaston, 1863: 219 N: CI (Tenerife)

excavatus lugubris Wollaston, 1864: 341 N: CI (Tenerife)

hupalupa Machado, 2007b: 16 N: CI (La Gomera)

hypogeus Machado, 2011a: 82 N: CI (El Hierro)

idafe García & Alonso-Zarazaga, 2011: 73 N: CI (La Palma)

inermis Machado, 2007b: 18 N: CI (La Gomera)

iruene Machado & García, 2010: 65 N: CI (La Palma)

laevis Roudier, 1957a: 18 N: CI (La Palma)

politus Roudier, 1957a: 19

oromii Machado, 2008b: 1282 N: CI (La Gomera)

sculptus Brullé, 1839: 71 (Otiorhynchus) N: CI (La Palma)

zarazagai subreflexus Machado & García, 2010: 68 N: CI (La Palma)

zarazagai zarazagai García & Oromí, 1997: 154 N: CI (La Palma)

Subgenus Mateuius Roudier, 1957a: 40 type species Lichenophagus auctus Wollaston, 1864

amplificatus Wollaston, 1865: 330 (Lichenophagus) N: CI (La Gomera)

auctus Wollaston, 1864: 363 (Lichenophagus) N: CI (El Hierro)

buccatrix Wollaston, 1865: Appendix, 54 (Lichenophagus) N: CI (La Gomera)

dilutus Machado, 2015: 105 N: CI (La Gomera)

merigensis Machado, 2015: 108 N: CI (La Gomera)

notatus Machado, 2015: 105 N: CI (La Gomera)

quadratus Machado, 2015: 108 N: CI (La Gomera)

teselinde Machado 2015: 106 N: CI (La Gomera)

Subgenus Pecoudius Roudier, 1957a type species Laparocerus cymbula Roudier, 1957a (= Cyphoscelis eliasenae Uyttenboogaart, 1929)

compactus Wollaston, 1864: 359 N: CI (Gran Canaria)

eliasenae Uyttenboogaart, 1929: 353 (Cyphoscelis) N: CI (Gran Canaria)

cymbula Roudier, 1957a: 38

franzi Machado, 2012: 10 N: CI (Gran Canaria)

sulcirostris Wollaston, 1864: 359 N: CI (Gran Canaria)

                  = canalirostris Marseul, 1872: 537

teldensis Machado, 2012: 11 N: CI (Gran Canaria)

[Pecoudius s.l.]

auarita Machado, 2016: 9 N: CI (La Palma)

bimbache Machado, 2011a: 74 N: CI (El Hierro)

brunneus Lindberg, 1953: 14 N: CI (Gran Canaria)

canescens Machado, 2016: 8 N: CI (Tenerife)

crassirostris Wollaston, 1864: 345 N: CI (Gran Canaria)

fraudulentus Machado, 2012: 14 N: CI (Gran Canaria)

freyi Uyttenboogaart, 1940: 675 N: CI (Tenerife)

garretai albosquamosus Machado, 2011b: 237 N: SG (Gran Pitón)

garretai garretai Uyttenboogaart, 1940: 56 N: SG (Selvagem Grande)

grayanus Wollaston, 1865: 309 (Atlantis) [RN] N: CI (Gran Canaria)

angustulus Wollaston, 1863: 219 (Atlantis) [HN]

hystricoides Machado, 2012: 16 N: CI (Gran Canaria)

inconspectus Roudier, 1957a: 27 N: CI (Gran Canaria)

lepidopterus lepidopterus Wollaston, 1864: 352 N: CI

              = bellopterus Marseul, 1872: 527

lepidopterus pecoudi Roudier, 1957a: 22 N: CI (Gran Canaria)

lopezi Machado, 2008b: 1285 N: CI (Gran Canaria)

microphthalmus Lindberg, 1950: 16 N: CI (Gran Canaria)

mulagua Machado, 2007b: 25 N: CI (La Gomera)

obsitus Wollaston, 1864: 361 N: CI (Gran Canaria)

persitus Marseul, 1872: 534

osorio Machado, 2012: 25 N: CI (Gran Canaria)

propinquus Lindberg, 1953: 14 N: CI (Gran Canaria)

punctiger Machado, 2016: 6 N: CI (Tenerife)

rugosivertex Machado, 2012: 18 N: CI (Gran Canaria)

semipilosus Machado, 2012: 23 N: CI (Gran Canaria)

separandus Lindberg, 1953: 14 N: CI (Gran Canaria)

soniae Machado, 2016: 35 N: CI (Gran Canaria)

tessellatus Brullé, 1839: 72 (Omias) N: CI (Tenerife)

tirajana Machado, 2012: 28 N: CI (Gran Canaria)

vicinus Lindberg, 1953: 15 N: CI (Gran Canaria)

Subgenus Pseudatlantis Machado, 2008a: 312 type species Atlantis excelsa Wollaston, 1854

abditus Roudier, 1963: 136 N: MR (Madeira)

excelsus Wollaston, 1854: 378 (Atlantis) N: MR (Madeira)

schaumii Wollaston, 1854: 379 (Atlantis) N: MR

forae Wollaston, 1854: 380 (Atlantis)

silvaticus Machado, 2008a: 319 N: MR (Madeira)

stuebeni Machado, 2008a: 321 N: MR (Madeira)

Subgenus Purpuranius Machado 2017: 33 type species Laparocerus maxorata Machado, 2011

calvus Machado, 2011b: 215 N: CI (Fuerteventura)

curvipes curvipes Lindberg, 1950: 17 N: CI (Tenerife)

curvipes espanoli Roudier, 1954: 84 N: CI (Fuerteventura, Lanzarote)

curvipes famarae Machado, 2011b: 219 N: CI (Lanzarote)

fraterculus Machado, 2011b: 211 N: CI (Fuerteventura)

longipennis Machado, 2011b: 213 N: CI (Fuerteventura)

maxorata Machado, 2011b: 208 N: CI (Fuerteventura)

Subgenus Wollastonius Machado, 2008a: 313 type species Omias ventrosus Wollaston, 1854

aenescens Wollaston, 1854: 383 (Omias) N: MR (Madeira)

angustulus Wollaston, 1857: 116 (Omias) N: MR (Madeira)

ventrosus Wollaston, 1854: 382 (Omias) N: MR (Madeira)

waterhousei Wollaston, 1854: 384 (Omias) N: MR


Genus Laparocerus, species incertae sedis

bacalladoi Machado, 2005: 540 N: CI (Tenerife)

depilis Roudier, 1957a: 22 N: CI (Tenerife)

ellipticus Wollaston, 1863: 220 N: CI

heres heres Machado, 2007b: 29 N: CI (La Gomera)

heres jocoensis Machado, 2007b: 35 N: CI (Tenerife)

inflatus Wollaston, 1865: Appendix, 41 N: CI (La Gomera)

tumens Marseul, 1872: 526

pilosiventris Machado, 2011a: 68 N: CI (La Gomera)

sanctaecrucis Machado, 2016: 11 N: CI (Tenerife)

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