Lizards, snails, caterpillars and beetles were common toys to play with when I was a child and enjoyed the countryside in La Palma or at the farm in Tenerife. Later, when I was a teenager I started a collection of beetles, and since then, I haven´t stopped, Entomology having become a Leitmotiv and umbilical cord with nature up to the present. Beetles were probably the hidden reason why I decided to study Biology at the University; when the pressure at home –a mild one– was to do something more “serious”, like Architecture, Agrarian engineering or Philosophy. However, I have never worked in Entomology professionally; but always as a happy and priviledged amateur.
When I finished my Biology Degree in La Laguna, the knowledge about Entomology of the Canary Islands was dispersed mainly in foreign institutions and journals. In 1976 I established, along with my wife and my brother, a cultural non-profit private association IBCER (International Bureau of Canarian Entomological Research) with the aim of collecting that information and offering a contact point for all researchers interested in our insect fauna.
At present, IBCER is dormant, but my library stays open and the same spirit of assistance to any colleague persists in the second floor at home, where I have established my cabinet (you can see a set of photos at the end of this page). Now, I have the intention to provide as much information as I can through this web-site, mainly on Coleoptera, bibliography, and whatever is in my hands, with the only limit being that of available time.
When you are young and looking at bugs, each group has attractive aspects: ants, springtails, spiders, grasshoppers, dragonflies… My interest in ground-beetles goes back to 1969, when I discovered a few black Microlestes close to my golf ball. Its perfect design just fascinated me. The choice was made and it received full support of Don José María Fernández, curator of the Museum of Natural History at Santa Cruz, whom I started to visit regularly.
I studied some 23.000 specimens from my own collection as well as from several museum collections (Madrid, Oxford, London, Paris, etc.). A total of 232 species live in the Canary Islands, 70% of which are endemic to the archipelago. I revised all the endemic taxa and described 1 tribe, 4 new genera and 29 new species. In parallel, I established 28 new synonyms. The Monografía de los carábidos de las islas Canarias, was published a year later (1992). It has 734 pages and 313 figures. Some chapters deal with aspects like phenology, dispersal power, and parasites, with some emphasis on biogeography and comments on evolution. But at that time, I did not perform phylogenetic analysis. That came later, but not with carabids.
You can access the Monograph here (PDF) or from the section Publication /Scientific, were all other papers dealing with Carabidae are available.
Having grown after the monograph on Carabidae, my interest in understanding how species form and why, islands are the best places to study evolution, and beetles the best candidates. I defined a profile for the ideal group: it has to be monophyletic, with as many species as possible, all endemic and without flying capacity. Winner by a wide margin: the genus Laparocerus Schönherr, 1834 (Curculionidae: Entiminae).
After 15 years of studying the genus I am still not finished, but getting closer to the end. The Laparocerus are not as rare as previously thought. They are easy to be found by night. I developed efficient techniques for sampling, having collected more than 32.000 specimens, many of them belonging to species previously unknown to science. I prospected the Azores, Madeira and Canary Islands consciously, repeating expeditions to the same island, or to the same place whenever needed. Additionally, I have studied the material kept in all major collections in European museums, plus that sent by colleagues. If you have a recalcitrant species of Laparocerus to be identified, please do not hesitate to write to me.
In the section of Production/Scientific there are several articles about Laparocerus, many describing new species. The number of species level taxa has increased from 123 (when I started) to 215, and there are some more in the pipeline.
I faced the phylogenetic study in collaboration with Dr. Mariano Hernández Ferrer of the University Institute of Tropical Diseases and Public Health of the Canary Islands, and I contracted technical assistance for the laboratory work. I financed the costs of the initial molecular analyses with a grant from the Fundación Biodiversidad (Madrid), which I obtained in 2004-2005. Obviously, I had to update my notions of Genetics and deepen my knowledge of new methods of inferring phylogenies, as well as familiarise myself with a complex arsenal of computer programmes. Later, I had to extend the molecular analyses to more markers and more species, this time with the collaboration of IPNA (CSIC) and hiring more assistants.
In the Production/Scientific section are my articles on Laparocerus, including the descriptions of new species published progressively. The number of taxa at the species level has increased from 123 (when I started) to 265. Finally, the monograph that gathers all the information generated in the last twenty years, I wrote it directly in English and I made the layout taking advantage of the COVID19 pandemic period. It was published in December 2022, in large format,
Since Coleoptera Atlantidum of Wollaston (1865), by the end of the XX Century, there was no modern account of beetles from the Canary Islands. In collaboration with Dr. Pedro Oromí, from the University of La Laguna, and with the assistance of many colleagues, we published a new catalogue in 2000 (see PDF), updating to 2061 the total number of species and subspecies, of which 1247 are endemic. We also registered 325 recognized synonyms.
When I find a suitable database and some time, it is my intention to include the Canarian Catalogue in this website for online access. For the moment, I keep it in my desktop running on Procite, and oldie, but performing well.
One of my main goals at IBCER was to find all the entomological bibliography related to the Canary Islands. That was in the times of cards, typing machines, carbon copies, ordinary letters and smailmail to contact other colleagues, visiting libraries in person, etcetera; something unimaginable for the Internet generation. In 1976 I finished a preliminary list, and in 1987 the book A Bibliography of the Canary Island Entomology was published, which was a milestone.
After finishing the monograph on Laparocerus in 2022, I thought about the best use I could make out of me and the time available after my retirement. As taxonomists are becoming something of an endangered species, I decided to apply myself to the elaboration of keys for the identification of the Canarian Coleoptera. We are talking about 2500 species distributed in 71 families. Except for carabids and curculionids, nothing similar exists. Only a few dispersed genus keys and rarely in Spanish
Needless to say, I am grateful in advance for any feedback: warnings of errors, absences, suggestions, incorrect images, etc. Please write directly to antonio.machado@telefonica.net.
CABINET WORK
COLLABORATORS
SAMPLING TECHNIQUES
At present, the application for collecting species must be submitted electronically and in Spanish to the Biodiversity Service of the Canary Islands Government. In addition, permission must then be obtained from each island council –also in Spanish and electronically (not always)– to collect in protected areas, if that is the case, and the councils do not make a decision without first receiving the resolution from the Canary Islands Government. Fortunately, the latter usually sends a copy of its resolution to the island council (Cabildo insular) concerned. I am afraid that a bureaucratic maquis has been created, to be overcome with patience, perseverance, and the necessary foresight. Good luck!
Genera
Subgenera
Species
Subspecies
Updated Dec. 2023
Subgenus Amyntas Wollaston, 1865: Appendix, 55 type species Lichenophagus incomptus Wollaston, 1865
Canopus Wollaston, 1864: 337 [HN] type species Atlantis subnebulosa Wollaston, 1864
arrochai Machado, 2009b: 190 N: CI (La Palma)
bellus Roudier, 1957a: 31 N: CI (Tenerife)
cristatus Machado, 2009b: 193 N: CI (La Palma)
fernandezi Roudier, 1957a: 34 N: CI
incomptus Wollaston, 1865: Appendix, 55 (Lichenophagus) N: CI (El Hierro)
ruteri Roudier, 1957a: 33 N: CI (Tenerife)
subnebulosus Wollaston, 1864: 337 (Atlantis) N: CI (Gran Canaria)
tanausu Machado, 2009b: 195 N: CI (La Palma)
tetricus Boheman, 1834b: 542 (Eremnus) N: CI (Tenerife)
simplex Brullé, 1839: 71 (Otiorhynchus)
tibialis isorae Machado, 2021: 148 N: CI (Tenerife)
tibialis tibialis Wollaston, 1864: 338 (Atlantis) N: CI (Tenerife)
uyttenboogaarti Zumpt, 1940: 677 N: CI (Tenerife)
Subgenus Anillobius Fauvel, 1907: 71 type species Anillobius solifuga Fauvel, 1907
portosantoi Franz, 1970: 3 (Anillobius) N: MR (Porto Santo)
rhizophilus Machado, 2021: 142 N: MR (Madeira)
solifuga Fauvel, 1907: 72 (Anillobius) N: MR (Madeira)
Subgenus Aridotrox Machado, 2017: 33 type species Laparocerus rasus Wollaston, 1864
chinijo Machado, 2021: 144 N: CI (La Graciosa, Montaña Clara)
colonnellii Machado, 2011b: 232 N: CI (Fuerteventura)
dispar Wollaston, 1864: 357 N: CI (Lanzarote)
inexpectatus Machado, 2011b: 229 N: MO
tessellatus Hustache, 1932b: 43 (Cyclomaurus) [HN]
rasus betancor Machado, 2011b: 223 N: CI (Fuerteventura)
rasus jandiensis Machado, 2011b: 224 N: CI (Fuerteventura)
rasus rasus Wollaston, 1864: 354 N: CI (Lanzarote, Montaña Clara)
susicus montanus Machado, 2011b: 229 N: MO
susicus susicus Escalera, 1914: 424 (Cyclobarus) N: MO
xericola Machado, 2011b: 231 N: CI (Fuerteventura)
Subgenus Atlantis Wollaston, 1854b: 361 type species Atlantis lamellipes Wollaston, 1854
calcatrix Wollaston, 1854b: 366 (Atlantis) N: MR (Madeira)
calcator Marseul, 1873b: 546
clavatus Wollaston, 1854b: 363 (Atlantis) N: MR (Madeira)
lamellipes hobbit Machado, 2008a: 318 N: MR (Madeira)
lamellipes lamellipes Wollaston, 1854b: 364 (Atlantis) N: MR (Madeira)
noctivagans lauripotens Wollaston, 1854b: 369 (Atlantis) N: MR (Madeira)
australis Wollaston, 1864: 370 (Atlantis)
lindbergi Roudier, 1963b: 133 N: MR (Madeira)
madeirensis Machado, 2008a: 316 N: MR (Madeira)
noctivagans lauripotens Wollaston, 1854b: 369 (Atlantis) N: MR (Madeira)
noctivagans noctivagans Wollaston, 1854b: 367 (Atlantis) [NP] N: MR (Madeira)
piceus Boheman, 1834b: 531 [NO]
serrado Machado, 2008a: 319 N: MR (Madeira)
undulatus Wollaston, 1862c: 289 N: MR (Madeira)
vespertinus Wollaston, 1854b: 371 (Atlantis) N: MR (Madeira)
Subgenus Atlantodes Machado, 2008a: 312 type species Atlantis mendax Wollaston, 1854
colasi Roudier, 1958b: 205 N: MR (Madeira)
erberi Machado, 2021: 142 N: MR (Deserta Grande)
inconstans Wollaston, 1854b: 375 (Atlantis) N: MR (Porto Santo)
instabilis Wollaston, 1854b: 377 (Atlantis) N: MR (Porto Santo)
lanatus Wollaston, 1854b: 372 (Atlantis) N: MR (Madeira)
mendax Wollaston, 1854b: 376 (Atlantis) N: MR (Porto Santo)
navicularis Wollaston, 1854b: 374 (Atlantis) N: MR (Porto Santo)
prainha Machado, 2008a: 323 N: MR (Madeira)
Subgenus Belicarius Machado, 2017: 35 type species Laparocerus mendicus Wollaston, 1864
bentejui bentejui Machado, 2012: 7 N: CI (Gran Canaria)
bentejui delicatulus Machado, 2012: 8 N: CI (Gran Canaria)
bentejui robustus Machado, 2012: 10 N: CI (Gran Canaria)
exiguus Machado, 2007b: 11 N: CI (La Gomera)
exophthalmus cisti Machado, 2021: 152 N: CI (La Gomera)
exophthalmus exophthalmus Machado, 2007b: 12 N: CI (La Gomera)
feloi Machado, 2009b: 203 N: CI (La Palma)
longiclava Har. Lindberg, 1953: 15 N: CI
magnificus Machado, 2011a: 70 N: CI
mateui mateui Roudier, 1954d: 85 N: CI (La Gomera)
mateui tuberosus Machado, 2011a: 74 N: CI (El Hierro)
mendicus Wollaston, 1864: 355 N: CI (El Hierro)
morrisi Machado, 2009b: 218 N: CI (La Palma)
oculatissimus Machado, 2007b: 14 N: CI (La Gomera)
rotundatus Machado, 2011a: 65 N: CI (La Gomera)
sanchezi arures Machado, 2016: 33 N: CI (La Gomera)
sanchezi sanchezi Roudier, 1957a: 24 N: CI (El Hierro)
nitidirostris Roudier, 1957a: 25
tarsalis Machado, 2009b: 205 N: CI (La Palma)
Subgenus Bencomius Machado, 2017: 35 type species Laparocerus grossepunctatus Wollaston, 1864
bolivari Uyttenboogaart, 1937: 97 N: CI (Tenerife)
boticarius Machado, 2007a: 39 N: CI (Tenerife)
combrecitensis Roudier, 1957c: 90 [RN] N: CI (La Palma)
palmensis Roudier, 1957a: 20 [HN]
crassifrons Wollaston, 1864: 345 N: CI (Tenerife)
edaphicus Machado, 2008b: 1278 N: CI (Tenerife)
escaleraorum Uyttenboogaart, 1937: 96 N: CI (Tenerife)
escalerai: Uyttenboogaart, 1937: 96 [IOS, NA]
gomerensis Har. Lindberg, 1953: 15 N: CI (La Gomera)
grossepunctatus Wollaston, 1864: 344 N: CI (Tenerife)
scapularis Wollaston, 1864: 347 N: CI (Tenerife)
wollastoni Marseul, 1873b: 521
subparallelus Machado, 2007a: 43 N: CI (Tenerife)
supranubius Machado, 2009b: 210 N: CI (La Palma)
tenuepunctatus commixtus Machado, 2021: 151 N: CI (Tenerife)
tenuepunctatus oppositus Machado, 2016: 16 N: CI (Tenerife)
tenuepunctatus tenuepunctatus Roudier, 1957a: 20 N: CI (Tenerife)
undatus Wollaston, 1864: 342 N: CI (Tenerife)
Subgenus Bentaygus Machado, 2022: 469 type species: Laparocerus vicinus Lindberg, 1953
brunneus Har. Lindberg, 1953: 14 N: CI (Gran Canaria)
propinquus Har. Lindberg, 1953: 14 N: CI (Gran Canaria)
vicinus Har. Lindberg, 1953: 15 N: CI (Gran Canaria)
Subgenus Canariotrox Machado, 2017: 35 type species Laparocerus inaequalis Wollaston, 1864
abona Machado, 2016: 14 N: CI (Tenerife)
acyphus Machado, 2009b: 189 N: CI (La Palma)
aeneotinctus aeneotinctus Machado, 2009b: 185 N: CI (La Palma)
aeneotinctus femoralis Machado, 2009b: 188 N: CI (La Palma)
aguiari Machado, 2007a: 36 N: CI (Tenerife)
crassus Roudier, 1957a: 28 N: CI (Tenerife)
estevezi Machado, 2012: 13 N: CI (Gran Canaria)
inaequalis globulipennis Wollaston, 1864: 349 N: CI (La Palma)
globosipennis Marseul, 1873b: 531
inaequalis hirtus Wollaston, 1864: 348 N: CI (Gran Canaria)
doramasensis Uyttenboogaart, 1929b: 352
inaequalis inaequalis Wollaston, 1863: 220 N: CI (Tenerife)
occidentalis Wollaston, 1864: 350 N: CI (El Hierro)
rugosicollis Uyttenboogaart, 1937: 98 N: CI (Tenerife)
tauce Machado, 2016: 16 N: CI (Tenerife)
vestitus Wollaston, 1864: 358 N: CI
velatus Marseul, 1873b: 532,
affinis Wollaston, 1864: 358
Subgenus Doramsasius Machado, 2022: 474 type species Atlantis grayana Wollaston, 1865
fraudulentus Machado, 2012: 14 N: CI (Gran Canaria)
grayanus Wollaston, 1865: 309 (Atlantis) [RN] N: CI (Gran Canaria)
angustulus Wollaston, 1863: 219 (Atlantis) [HN]
hystricoides Machado, 2012: 16 N: CI
inconspectus Roudier, 1957a: 27 N: CI (Gran Canaria)
semipilosus Machado, 2012: 23 N: CI (Gran Canaria)
Subgenus Faycanius Machado, 2012: 29 type species Otiorhynchus squamosus Brullé, 1839
alluaudi Uyttenboogaart, 1940a: 55 N: CI (Gran Canaria)
anniversarius Machado, 2012: 37 N: CI (Gran Canaria)
arcanus Machado, 2012: 32 N: CI (Gran Canaria)
aytamis Machado, 2012: 33 N: CI (Gran Canaria)
canutus Machado, 2021: 144 N: CI (Gran Canaria)
dissidens dissidens Machado, 2012: 35 N: CI (Gran Canaria)
dissidens spolonifer Machado, 2021: 146 N: CI (Gran Canaria)
reptans Machado, 2021: 146 N: CI (Gran Canaria)
squamosus Brullé, 1839: 71 (Otiorhynchus) N: CI (Gran Canaria)
seniculus Wollaston, 1864: 353
tasarticus Machado, 2012: 21 N: CI (Gran Canaria)
vallei Machado, 2021: 144 N: CI (Gran Canaria)
Subgenus Fernandezius Roudier, 1957a: 40 type species Lichenophagus tesserula Wollaston, 1864
anagae Machado, 2015: 126 N: CI (Tenerife)
anisomorphus Machado, 2021: 149 N: CI (Tenerife)
campestris campestris Machado, 2015: 132 N: CI (Tenerife)
campestris garafianus Machado, 2021: 151 N: CI (La Palma)
hadrocerus Machado, 2021: 149 N: CI (Tenerife)
impressicollis Wollaston, 1864: 368 (Lichenophagus) N: CI (Tenerife)
× intermedius Roudier, 1957a: 43 (Lichenophagus) N: CI (Tenerife)
macilentus Machado, 2015: 125 N: CI (Tenerife)
persimilis Wollaston, 1864: 365 (Lichenophagus) N: CI (Tenerife)
pitys Machado, 2021: 149 N: CI (La Palma)
sculptipennis montivagans Machado, 2013: 69 [RN] N: CI (La Palma)
palmensis Roudier, 1957a: 43 (Lichenophagus) [HN]
sculptipennis sculptipennis Wollaston, 1864: 367 (Lichenophagus) N: CI (La Palma)
seriesetosus Wollaston, 1864: 365 (Lichenophagus) N: CI (La Palma)
sonchiphagus Machado, 2015: 120 N: CI (Tenerife)
subcalvus Wollaston, 1864: 336 (Lichenophagus) N: CI (El Hierro)
subnodosus Wollaston, 1864: 336 (Lichenophagus) N: CI (Tenerife)
tenicola Machado, 2015: 128 N: CI (Tenerife)
tesserula Wollaston, 1864: 364 (Lichenophagus) N: CI (Tenerife)
Subgenus Fortunotrox Machado, 2011a: 58 type species Laparocerus gracilis Wollaston, 1865
acutipennis Machado, 2007b: 5 N: CI (La Gomera)
benchijigua Machado, 2007b: 6 N: CI (La Gomera)
confusus Machado, 2011a: 62 N: CI (La Gomera)
crotchi Machado, 2016: 30 N: CI (La Gomera)
decipiens Machado, 2009b: 214 N: CI (La Palma)
depressus Machado, 2007b: 7 N: CI (La Gomera)
gracilis Wollaston, 1864: 356 N: CI (La Gomera)
gomeraecola Marseul, 1873b: 531
idafe García & Alonso-Zarazaga, 2011: 73 N: CI (La Palma)
indutus Wollaston, 1865: Appendix, 53 N: CI (La Gomera)
tectus Marseul, 1873b: 535
juelensis eremita Machado, 2021: 147 N: CI (La Gomera)
juelensis juelensis Machado, 2011a: 64 N: CI (La Gomera)
orone aridane Machado, 2009b: 211 N: CI (La Palma)
orone hierrensis Machado, 2011a: 80 N: CI (El Hierro)
orone orone Machado, 2007b: 3 N: CI (La Gomera)
puncticollis Wollaston, 1864: 362 N: CI (El Hierro)
scitulus Machado, 2021: 146 N: CI (La Gomera)
spinimanus Machado, 2007b: 9 N: CI (La Gomera)
subopacus divergens Machado, 2021: 147 N: CI (La Gomera)
subopacus subopacus Wollaston, 1865: Appendix, 52 N: CI (La Gomera)
Subgenus Guanchotrox Alonso-Zarazaga & Lyal, 1999: 11 [RN] type species Laparocerus canariensis Boheman, 1843
Amphora Wollaston, 1864: 335 [HN] type species Laparocerus canariensis Boheman, 1843
amicorum Machado, 2009b: 212 N: CI (La Palma)
astralis Machado, 2009b: 206 N: CI (La Palma)
canariensis Boheman, 1842b: 228 N: CI (Tenerife)
cephalotes Machado, 2011a: 78 N: CI (El Hierro)
debilis Wollaston, 1865: Appendix, 53 N: CI (Tenerife)
dissimilis alticola Machado, 2016: 22 N: CI (Tenerife)
dissimilis dissimilis Har. Lindberg, 1950: 17 N: CI (Tenerife)
dissimilis infernalis Machado, 2016: 23 N: CI (Tenerife)
elongatus denudatus Machado, 2021: 151 N: CI (La Palma)
elongatus elongatus Machado, 2009b: 198 N: CI (La Palma)
elongatus mucronatus Machado, 2009b: 201 N: CI (La Palma)
gerodes Machado, 2016: 23 N: CI (La Gomera)
humeralis Machado, 2007b: 23 N: CI (La Gomera)
junonius Machado, 2007b: 19 N: CI (La Gomera)
marmoratus Machado, 2012: 22 N: CI (Gran Canaria)
obscurus daute Machado, 2016: 21 N: CI (Tenerife)
obscurus obscurus Wollaston, 1864: 355 N: CI (Tenerife)
obscurellus Marseul, 1873b: 529
obtriangularis Wollaston, 1864: 351 N: CI (Tenerife)
ornatus Machado, 2012: 20 N: CI (Gran Canaria)
roudieri Machado, 2007b: 22 N: CI (La Gomera)
tafadensis Machado, 2016: 28 N: CI (Tenerife)
tinguaro tabornoi Machado, 2016: 29 N: CI (Tenerife)
tinguaro tinguaro Machado, 2007a: 44 N: CI (Tenerife)
tirmensis Machado, 2012: 21 N: CI (Gran Canaria)
Subgenus Guayre Machado, 2022: 495 type species Omias tessellatus Brullé, 1839
auarita Machado, 2016: 9 N: CI (La Palma)
bimbache Machado, 2011a: 74 N: CI (El Hierro)
freyi canescens Machado, 2016: 8 N: CI (Tenerife)
freyi freyi Uyttenboogaart, 1940c: 675 N: CI (Tenerife)
freyi punctiger Machado, 2016: 6 N: CI (Tenerife)
freyi vicarius Machado, 2021: 152 N: CI (Tenerife)
garretai albosquamosus Machado, 2011b: 237 N: SG (Selvagem Pequena)
garretai garretai Uyttenboogaart, 1940a: 56 N: SG (Selvagem Grande)
microphthalmus Har. Lindberg, 1950: 16 N: CI (Gran Canaria)
obsitus Wollaston, 1864: 361 N: CI (Gran Canaria)
persitus Marseul, 1873b: 534
osorio Machado, 2012: 25 N: CI (Gran Canaria)
rugosivertex Machado, 2012: 18 N: CI (Gran Canaria)
tessellatus Brullé, 1839: 72 (Omias) N: CI (Tenerife)
tirajana Machado, 2012: 28 N: CI (Gran Canaria)
Subgenus Laparocerus Schoenherr, 1834b: 530 type species Laparocerus morio Boheman, 1834
Cyphoscelis Wollaston, 1854b: 356 type species Cyphoscelis distorta Wollaston, 1854
chaoensis cevadae Roudier, 1961a: 62 N: MR (Ilheu do Desembarcadouro)
chaoensis chaoensis Uyttenboogaart, 1940a: 57 N: MR (Ilheu de Chão, Bugio)
vandeli Roudier, 1958b: 199
cryptus Machado, 2008a: 315 N: MR
distortus Wollaston, 1854b: 357 (Cyphoscelis) N: MR (Madeira)
morio Boheman, 1834b: 531 N: MR (Madeira)
obesulus Desbrochers des Loges, 1904a: 63
Subgenus Lichenophagus Wollaston, 1854b: 389 type species Lichenophagus fritillus Wollaston, 1854
acuminatus Wollaston, 1854b: 391 N: MR (Deserta Grande)
fritillus Wollaston, 1854b: 390 N: MR (Porto Santo)
Subgenus Machadotrox Alonso-Zarazaga & Lyal, 1999: 10 type species Otiorhynchus sculptus Brullé, 1839
Wollastonia Uyttenboogaart, 1936: 255 (non Heer, 1852, nec Horn, 1873)
Wollastonicerus Uyttenboogaart, 1937: 96
aethiops aethiops Wollaston, 1864: 347 N: CI (El Hierro)
aethiops garajonay Machado, 2007b: 15 N: CI (La Gomera)
cavernarius Machado, 2011a: 84 N: CI (El Hierro)
chasnensis Machado, 2007a: 41 N: CI (Tenerife)
dacilae García, 1998: 45 N: CI (La Palma)
excavatus excavatus Wollaston, 1863: 219 N: CI (Tenerife)
lugubris Wollaston, 1864: 341
excavatus prosenemus Machado, 2021: 147 N: CI (La Palma)
hupalupa furtivus Machado, 2021: 147 N: CI (La Gomera)
hupalupa hupalupa Machado, 2007b: 16 N: CI (La Gomera)
hypogeus Machado, 2011a: 82 N: CI (El Hierro)
inermis inermis Machado, 2007b: 18 N: CI (La Gomera)
inermis majonae Machado, 2021: 148 N: CI (La Gomera)
iruene Machado & García, 2010: 65 N: CI (La Palma) laevis Roudier, 1957a: 18 N: CI (La Palma)
politus Roudier, 1957a: 19
machadoi García & González, 2006: 171 N: CI (La Palma)
oromii Machado, 2008b: 1282 N: CI (La Gomera)
sculptus Brullé, 1839: 71 (Otiorhynchus) N: CI (La Palma)
zarazagai criniger Machado, 2021: 148 N: CI (La Palma)
zarazagai subreflexus Machado & García, 2010: 68 N: CI (La Palma)
zarazagai zarazagai García & Oromí, 1997: 154 N: CI (La Palma)
Subgenus Mateuius Roudier, 1957a: 40 type species Lichenophagus auctus Wollaston, 1864
amplificatus Wollaston, 1865: 330 (Lichenophagus) N: CI (La Gomera)
auctus Wollaston, 1864: 363 (Lichenophagus) N: CI (El Hierro)
buccatrix Wollaston, 1865: Appendix, 54 (Lichenophagus) N: CI (La Gomera)
dilutus Machado, 2015: 105 N: CI (La Gomera)
merigensis Machado, 2015: 108 N: CI (La Gomera)
notatus Machado, 2015: 105 N: CI (La Gomera)
quadratus Machado, 2015: 108 N: CI (La Gomera)
teselinde Machado, 2015: 106 N: CI (La Gomera)
Subgenus Nanotrox Machado, 2022: 462 type species Laparocerus transversus Lindberg, 1950
buenavistae Roudier, 1957a: 29 N: CI (Tenerife)
palmensis Har. Lindberg, 1953: 16 N: CI (La Palma)
seminitens Har. Lindberg, 1950: 17 N: CI (Tenerife)
tenellus Wollaston, 1864: 362 N: CI (Tenerife)
transversus Har. Lindberg, 1950: 16 N: CI (Tenerife)
Subgenus Pecoudius Roudier, 1957a: 38 type species Laparocerus cymbula Roudier, 1957a (= Cyphoscelis eliasenae Uyttenboogaart, 1929)
compactus Wollaston, 1864: 359 N: CI (Gran Canaria)
crassirostris Wollaston, 1864: 345 N: CI (Gran Canaria)
eliasenae Uyttenboogaart, 1929b: 353 (Cyphoscelis) N: CI (Gran Canaria)
cymbula Roudier, 1957a: 38
franzi Machado, 2012: 10 N: CI (Gran Canaria)
sulcirostris Wollaston, 1864: 359 N: CI (Gran Canaria)
canalirostris Marseul, 1873b: 537
teldensis Machado, 2012: 11 N: CI (Gran Canaria)
Subgenus Proteotros Machado, 2022: 481 type species Laparocerus lepidopterus Wollaston, 1864
federico Machado, 2018: 11 N: CI (Gran Canaria)
lepidopterus lepidopterus Wollaston, 1864: 352 N: CI
bellopterus Marseul, 1873b: 527
lepidopterus pecoudi Roudier, 1957a: 22 N: CI (Gran Canaria)
lopezi Machado, 2008b: 1285 N: CI (Gran Canaria)
moyanus Machado, 2021: 152 N: CI (Gran Canaria)
mulagua Machado, 2007b: 25 N: CI (La Gomera)
separandus Har. Lindberg, 1953: 14 N: CI (Gran Canaria)
soniae Machado, 2016: 35 N: CI (Gran Canaria)
Subgenus Pseudatlantis Machado, 2008a: 312 type species Atlantis excelsa Wollaston, 1854
abditus Roudier, 1963b: 136 N: MR (Madeira)
excelsus Wollaston, 1854b: 378 (Atlantis) N: MR (Madeira)
schaumii forae Wollaston, 1854b: 380 (Atlantis)
schaumii schaumii Wollaston, 1854b: 379 (Atlantis) N: MR
silvaticus Machado, 2008a: 319 N: MR (Madeira)
stuebeni Machado, 2008a: 321 N: MR (Madeira)
Subgenus Purpuranius Machado, 2017: 33 type species Laparocerus maxorata Machado, 2011
calvus Machado, 2011b: 215 N: CI (Fuerteventura)
curvipes curvipes Har. Lindberg, 1950: 17 N: CI (Tenerife)
curvipes espanoli Roudier, 1954d: 84 N: CI (Fuerteventura, Lanzarote)
curvipes famarae Machado, 2011b: 219 N: CI (Lanzarote)
fraterculus Machado, 2011b: 211 N: CI (Fuerteventura)
longipennis Machado, 2011b: 213 N: CI (Fuerteventura)
maxorata Machado, 2011b: 208 N: CI (Fuerteventura)
Subgenus Wollastonius Machado, 2008a: 313 type species Omias ventrosus Wollaston, 1854
aenescens Wollaston, 1854b: 383 (Omias) N: MR (Madeira)
angustulus Wollaston, 1857: 116 (Omias) N: MR (Madeira)
desertarum Machado, 2021: 142 N: MR (Deserta Grande)
ventrosus Wollaston, 1854b: 382 (Omias) N: MR (Madeira)
waterhousei Wollaston, 1854b: 384 (Omias) N: MR
Genus Laparocerus, species incertae sedis
bacalladoi Machado, 2005: 540 N: CI (Tenerife)
depilis Roudier, 1957a: 22 N: CI (Tenerife)
ellipticus Wollaston, 1863: 220 N: CI
heres heres Machado, 2007b: 29 N: CI (La Gomera)
heres jocoensis Machado, 2007b: 35 N: CI (Tenerife)
inflatus Wollaston, 1865: Appendix, 41 N: CI (La Gomera)
tumens Marseul, 1873b: 526
pilosiventris Machado, 2011a: 68 N: CI (La Gomera)
sanctaecrucis Machado, 2016: 11 N: CI (Tenerife)
Update (January 2024) of genus Laparocerus in Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea (2nd Edition, 2017). Abbreviations: CI = Canary Islands, [HN] = homonym, [NO] = nomen oblitum, MR = Madeira archipelago, N = North Africa,
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Machado, A. 2007a: New species of Laparocerus Schönherr, 1834 (Coleoptera, Curculionidae) from the island of Tenerife, Canary Islands. Zootaxa 1457: 35-48.
Machado, A. 2007b: New species of Laparocerus Schönherr, 1834 from La Gomera, Canary Islands (Coleoptera, Curculionidae, Entiminae). Zootaxa 1643: 1-38.
Machado A. 2008a: New taxa of Laparocerus Schönherr, 1832 from Madeira and Porto Santo,with a key to the species of the Madeiran Archipelago (Coleoptera, Curculionidae, Entiminae). Graellsia 64 (2): 307-328.
Machado A. 2008b: Three new endogean species of Laparocerus Schönherr, 1834 from the Canary Islands (Coleoptera, Curculionidae). Journal of Natural History 43 (17-18): 1277-1288.
Machado A. 2009: Nuevos Laparocerus Schoenherr, 1934 de La Palma, islas Canarias (Coleoptera, Curculionidae, Entiminae). Graellsia 65 (2): 183-224.
Machado, A. 2011a. Nuevos taxones de Laparocerus Schoenherr, 1834 de El Hierro y La Gomera, islas Canarias (Coleoptera, Curculionidae, Entiminiae). Graellsia 67(1): 57-90.
Machado, A. 2011b. Los Laparocerus Schoenherr, 1834 de las Canarias orientales, islas Salvajes y Marruecos (Coleoptera, Curculionidae, Entiminae). Graellsia 67(2): 205-241.
Machado, A., 2012. Nuevos taxones de Laparocerus Schoenherr, 1834 de la isla de Gran Canaria (Coleoptera, Curculionidae, Entiminae). Boletín de la Sociedad Entomológica Aragonesa 51: 5-46.
Machado, A. 2013. Curculionidae: Entiminae: Laparocerini. Pp. 69-770, 288-292 in: Löbl I. & Smetana A. (eds.). Catalogue of Palaearctic coleoptera. Volume 8. Curculionoidea II. Stenstrup: Apollo Books.
Machado, A. 2015. Revisión de los antiguos Lichenophagus (= Laparocerus) de la Macaronesia (Coleoptera, Curculionidae, Entiminae). Boletín de la Sociedad Entomológica Aragonesa 57: 92-141.
Machado, A. 2016. Laparocerus Schoenherr, 1834 nuevos de Canarias y claves de las especies de la isla de Tenerife (Coleoptera, Curculionidae, Entiminae). Graellsia 72: e049.
Machado, A., 2017. Descriptions. In; Machado, A., Rodríguez-Expósito, E, López M, & Hernández M (2017) Phylogenetic analysis of the genus Laparocerus (Coleoptera, Curculionidae, Entiminae), with comments on colonization and diversification in Macaronesia. ZooKeys 651: 1-77. doi: 10.3897/zookeys.651.10097
Machado, A. 2017. Tribe Laparocerini Lacordaire, 1863. In: Alonso-Zarazaga M. A. (ed. Cooperative catalogue of Palaearctic Coleoptera. Sociedad Entomológica Aragonesa. Zaragoza: Pp. 280-285.
Machado, A. 2018. Laparocerus federico n. sp. nueva especie hipogea de la isla de Gran Canaria (Coleoptera, Curculionidae, Entiminae). Revista de la Academia Canaria de Ciencias 30: 9-18.
Machado, A. 2021. Additional new taxa of Laparocerus Schoenherr, 1834 from Macaronesia (Coleoptera, Curculionidae, Entiminae). Boletín de la Sociedad Entomológica Aragonesa 69: 141-157.
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